List of pterosaur-bearing stratigraphic units

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This is a list of stratigraphic units, where pterosaur fossils have been recovered from. Units listed are all either formation rank or higher (e.g. group).

Name Age Location Fossil species

Aguja Formation

Alamyshik Formation

Whitby Mudstone Formation

Amminadav Formation or Bet-Meir Formation

Antlers Formation

Apón Formation

Argiles d'Ecqueville

Argiles d'Octeville


Argilliti di Riva di Solto


Artoles Formation

Austin Group


?Bahia Series

Bakhar Svita

Balabansai Svita

Bissekty Formation


Blackhawk Formation

Blesa Formation

Blue Lias

Bostobin Formation

Aralazhdarcho, "Samrukia"[1]

Buda Limestone

Budden Canyon Formation

Calcaires tâchetés


Calizas de La Huergina Formation

Calizas y Margas de Sierra Perenchiza Formation

Camadas de Guimarota

Camarillas Formation

Cambridge Greensand


Cañadon Asfalto Formation

Candeleros Formation

Caturrita Formation[3]


Cerro del Pueblo Formation

Chenini Formation

Chico Formation

?Chinle Formation

Chipping Norton Formation

Chulec Formation

?Clarens Formation

Cornet Bauxite

Cotswold Slate

Crato Formation

Csehbánya Formation


Curtis/Stump Formation

Daohugou Beds

Jeholopterus, Pterorhynchus, Wukongopterus[4]

Dashanpu Formation


Densus-Ciula Formation


Dinosaur Park Formation


Dockum Group

Dolomia di Forni

Eudimorphodon, Preondactylus

Dunvegan Formation

Dzharakuduk Formation

Eagle Ford Formation

Ejinhoro Formation

El Castillar Formation

El Gallo Formation

Elligserbrink Shale

Enciso Group

Erlhaz Formation

Eumeralla Formation

Eutaw Formation

Exu Formation

Fleming Fjord Formation


Forest Marble Formation

Gault Clay


Gisement des Bessons


Glen Rose Formation

Gosau Formation

Gramme Formation


"Great Oolite"

Greenhorn Limestone

?Gres a Avicula contorta

Grès de la Crèche inférieurs

Hakobuchi Group

?"Hallau Bonened"

Hampen Marly Formation

Hanson Formation

Hasandong Formation

Hastings Beds

Coloborhynchus, Lonchodectes

Hekou Formation

Hell Creek Formation

Himenoura Group

Huachihuanhe Formation


Hudspeth Formation


Ilek Formation

Intertrappean Beds

Isalo III Formation

Jagua Formation

Cacibupteryx, Nesodactylus

Javelina Formation


Jiufotang Formation

Chaoyangopterus, Eoazhdarcho, Eopteranodon, "Huaxiapterus", Jidapterus, Liaoningopterus, Liaoxipterus, Nemicolopterus, Nurhachius, Sinopterus

Karabastau Formation

Batrachognathus, Sordes

Kayenta Formation


Kem Kem Beds

Alanqa,[6] Siroccopteryx[7]

Khilok Formation

Khodzhakul Formation

Kimmeridge Clay


Kössen beds

Kota Formation


Kuwajima Formation

La Amarga Formation

La Boca Formation

"Dimorphodon" weintraubi

La Cruz Formation


Lagarcito Formation


Lance Formation

Lastres Formation

Lezas Formation[9]


Lianmuqin Formation

Dsungaripterus, Lonchognathosaurus,[10] Noripterus

"Lithographic Limestones"

Logbadjeck Formation

Lohan Cura Formation

Lower Chalk

Lower Kimmeridgian Mudstones

Luohangdong Formation

Marne des Dives

Marnes Rouges de Roquelongue

Marnes Rouges Inférieures Formation

Marnes d'Auzas Formation

Melovatka Formation

Meng-Yin Formation

Merchantville Formation

Mesaverde Group

Middle Chalk

Middle Iser Shales

Mifune Group

Miria Formation

Mishash Formation

Mornsheimer Limestone

Altmuehlopterus, Germanodactylus, Pterodactylus, Ardeadactylus, Rhamphorhynchus

Mooreville Chalk

Morrison Formation

Mowry Shale

Murtoi Formation

Niobrara Chalk

Dawndraco,[11] Geosternbergia,[11] Nyctosaurus, Pteranodon

North Horn Formation

Nugget Sandstone


Nusplingen Limestone

Cycnorhamphus, Ardeadactylus, Rhamphorhynchus

Okurodani Formation

Oncala Formation

Öösh Formation

Oxford Clay Formation

Patuxent Formation

Paw Paw Formation


Pierre Shale


Portezuelo Formation

Posidonia Shale

Campylognathoides, Dorygnathus

Pudovinko Formation



Purbeck Beds

Gnathosaurus, Plataleorhynchus

Qingshan Formation

Qiqu Formation

Quebrada Monardes Formation


Reuchenette Formation

Río Belgrano Formation

Rybushka Formation

Bogolubovia,[12] Volgadraco[13]

Sannine Formation


Sânpetru Formation

Santa Ana Formation

Santana Formation


?Santa Rosa Formation

Seefelder Beds

Sekmenevka Formation

Sharp's Hill Formation

Shishugou Formation

?Sloan Canyon Formation

Solnhofen Plattenkalk

Altmuehlopterus, Anurognathus, Aurorazhdarcho,[16] Ctenochasma, Cycnorhamphus, Germanodactylus, Gnathosaurus, Pterodactylus, Ardeadactylus, Rhamphorhynchus, Scaphognathus

Solothurn Turtle Limestone

?Steinmergel Group

Stonesfield Slate

Summerville Formation

Sundance Formation

?"Syren Bonebed"

Tahora Formation

Tangshang Formation


Tarrant Formation


Taynton Limestone Formation

Tendaguru Formation


Tiaojishan Formation

Archaeoistiodactylus,[18] Changchengopterus,[19] Darwinopterus,[20] Fenghuangopterus[21] Jianchangopterus,[22] Jianchangnathus,[23] Qinglongopterus[24]

Tlayúa Formation

Toolebuc Formation

Aussiedraco,[25] Mythunga[26]

Toxaster Limestone

Tsagaantsav Svita


Turners Falls Formation

Two Medicine Formation


Uhangri Formation

"Unnamed unit of fissure fills"

"Unnamed unit of fissure fills"

Upper Chalk

Upper ‘couche III’, phosphate deposits


Upper Elliot Formation

Upper Greensand


Upper Sables de Glos

Vaca Muerta Formation


Vectis Formation



Wessex Formation


White Chalk

White Chalk Formation

"Wierzbica Oolite and Platy Limestones"

Yalovach Formation

Yezo Group

Yixian Formation

Beipiaopterus, Boreopterus, Cathayopterus, Dendrorhynchoides, Elanodactylus,[27] Eosipterus, Feilongus, Gegepterus, Gladocephaloideus,[28] Haopterus, Ningchengopterus,[29] Zhenyuanopterus[30]

Zhirkindek Formation

Zorzino Limestone

Eudimorphodon, Peteinosaurus, ?Preondactylus

Züünbayan Formation

See also


  1. ^ Buffetaut, E. "Samrukia nessovi, from the Late Cretaceous of Kazakhstan: A large pterosaur, not a giant bird". Annales de Paléontologie. 97: 133–138. doi:10.1016/j.annpal.2011.10.001.
  2. ^ Martill, David M.; Unwin, David M. (2011). "The world's largest toothed pterosaur, NHMUK R481, an incomplete rostrum of Coloborhynchus capito (Seeley 1870) from the Cambridge Greensand of England". Cretaceous Research. 34: 1–9. doi:10.1016/j.cretres.2011.09.003.
  3. ^ J. F. Bonaparte, C. L. Schultz, and M. B. Soares. 2010. Pterosauria from the Late Triassic of southern Brazil. In S. Bandyopadhyay (ed.), New Aspects of Mesozoic Biodiversity, Lecture Notes in Earth Sciences 132:63-71.
  4. ^ Wang X.; Kellner, A. W. A.; Jiang S.; Meng X. (2009). "An unusual long-tailed pterosaur with elongated neck from western Liaoning of China". Anais da Academia Brasileira de Ciências. 81 (4): 793–812. doi:10.1590/S0001-37652009000400016.
  5. ^ Robert M. Sullivan and Denver W. Fowler (2011). "Navajodactylus boerei, n. gen., n. sp., (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (upper Campanian) of New Mexico" (PDF). Fossil Record 3. New Mexico Museum of Natural History and Science, Bulletin. 53: 393–404.CS1 maint: Uses authors parameter (link)
  6. ^ Ibrahim N.; Unwin D.M.; Martill D.M.; Baidder L.; Zouhri S. (2010). "A New Pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco". PLoS ONE. 5 (5): e10875. Bibcode:2010PLoSO...510875I. doi:10.1371/journal.pone.0010875. PMC 2877115. PMID 20520782.
  7. ^ Rodrigues, T. and Kellner, A.W.A. (2008). "Review of the pterodactyloid pterosaur Colobrohynchus." Pp. 219–228 in: Hone, D.W.E. and Buffetaut, E. (eds), Flugsaurier: pterosaur papers in honour of Peter Wellnhofer. Zitteliana B, 28.
  8. ^ David M. Martill and Steve Etches (2011). "A new monofenestratan pterosaur from the Kimmeridge Clay Formation (Upper Jurassic, Kimmeridgian) of Dorset, England" (PDF). Acta Palaeontologica Polonica. in press. doi:10.4202/app.2011.0071.CS1 maint: Uses authors parameter (link)
  9. ^ Fuentes Vidarte, C., Meijide Calvo, M. (2010). "Un nuevo pterosaurio (Pterodactyloidea) en el Cretácico Inferior de La Rioja (España) (A new pterosaur (Pterodactyloidea) from the Lower Cretaceous of La Rioja (Spain))" (PDF). Boletín Geológico y Minero. 121 (3): 311–328. Archived from the original (PDF) on 2011-07-20.CS1 maint: Multiple names: authors list (link)
  10. ^ Maisch M.W.; Matzke A.T.; Sun Ge (2004). "A new dsungaripteroid pterosaur from the Lower Cretaceous of the southern Junggar Basin, north-west China". Cretaceous Research. 25: 625–634. doi:10.1016/j.cretres.2004.06.002.
  11. ^ a b Kellner, A.W.A. (2010). "Comments on the Pteranodontidae (Pterosauria, Pterodactyloidea) with the of two new species" (PDF). Anais da Academia Brasileira de Ciências. 82 (4): 1063–1084. doi:10.1590/S0001-37652010000400025.
  12. ^ Bogolubov, N.N. (1914). "O pozvonk’ pterodaktilya uz’ vyerkhnyem’lovyikh’ otlozhyenii Saratovskoi gubyernii. [On a pterodactyle vertebra from Upper Cretaceous deposits of the Government of Saratoff]." Annuaire geologique et mineralogique de la Russie, 16(1): 1-7. (Russian text, French resume).
  13. ^ Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (October 2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal. 42 (6): 634–642. doi:10.1134/S0031030108060099.
  14. ^ Ross A. Elgin & Eberhard Frey (2011). "A new azhdarchoid pterosaur from the Cenomanian (Late Cretaceous) of Lebanon". Swiss Journal of Geosciences. in press: 21–33. doi:10.1007/s00015-011-0081-1.
  15. ^ David M. Martill (2011). "A new pterodactyloid pterosaur from the Santana Formation (Cretaceous) of Brazil". Cretaceous Research. 32 (2): 236–243. doi:10.1016/j.cretres.2010.12.008.
  16. ^ Eberhard Frey, Christian A. Meyer and Helmut Tischlinger (2011). "The oldest azhdarchoid pterosaur from the Late Jurassic Solnhofen Limestone (Early Tithonian) of Southern Germany". Swiss Journal of Geosciences. 104: 35–55. doi:10.1007/s00015-011-0073-1.
  17. ^ Myers, Timothy S. (2010). "A new ornithocheirid pterosaur from the Upper Cretaceous (Cenomanian–Turonian) Eagle Ford Group of Texas" (pdf). Journal of Vertebrate Paleontology. 30 (1): 280–287. doi:10.1080/02724630903413099.
  18. ^ Lü Junchang and Fucha Xiaohui (2011). "A new pterosaur (Pterosauria) from Middle Jurassic Tiaojishan Formation of western Liaoning, China". Global Geology Z1: 113–118.
  19. ^ Lü, J. (2009). "A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China". Acta Geologica Sinica (English Edition). 83 (2): 189–199. doi:10.1111/j.1755-6724.2009.00062.x.
  20. ^ Lü, J.; Unwin, D.M.; Jin, X.; Liu, Y.; Ji, Q. (2010). "Evidence for modular evolution in a long-tailed pterosaur with a pterodactyloid skull". Proceedings of the Royal Society B. 277 (1680): 383–389. doi:10.1098/rspb.2009.1603. PMC 2842655. PMID 19828548.
  21. ^ Lu, J.; Fucha, X.; Chen, J. (2010). "A new scaphognathine pterosaur from the Middle Jurassic of western Liaoning, China". Acta Geoscientica Sinica. 31 (2): 263–266.
  22. ^ Lü Junchang & Bo Xue (2011). "A New Rhamphorhynchid Pterosaur (Pterosauria) from the Middle Jurassic Tiaojishan Formation of Western Liaoning, China". Acta Geologica Sinica. 85 (5): 977–983. doi:10.1111/j.1755-6724.2011.00531.x.
  23. ^ Xin Cheng; Xiaolin Wang; Shunxing Jiang; Alexander W.A. Kellner (2012). "A new scaphognathid pterosaur from western Liaoning, China". Historical Biology. 24: 101–111. doi:10.1080/08912963.2011.635423.
  24. ^ Lü, J., Unwin, D.M., Zhao, B., Gao, C. and Shen, C. (2012). "A new rhamphorhynchid (Pterosauria: Rhamphorhynchidae) from the Middle/Upper Jurassic of Qinglong, Hebei Province, China" (PDF). Zootaxa. 3158: 1–19.CS1 maint: Multiple names: authors list (link)
  25. ^ Kellner, Alexander W.A.; Taissa Rodrigues; Fabiana R. Costa (2011). "Short note on a pteranodontoid pterosaur (Pterodactyloidea) from western Queensland, Australia" (PDF). Anais da Academia Brasileira de Ciências. 83 (1): 301–308. doi:10.1590/S0001-37652011000100018.
  26. ^ Molnar, Ralph E.; Thulborn, R.A. (2008). "An incomplete pterosaur skull from the Cretaceous of north-central Queensland, Australia". Arquivos do Museu Nacional, Rio de Janeiro. 65 (4): 461–470.
  27. ^ Andres, B.; Ji Q. (2008). "A new pterosaur from the Liaoning Province of China, the phylogeny of the Pterodactyloidea, and convergence in their cervical vertebrae". Palaeontology. 51 (2): 453–469. doi:10.1111/j.1475-4983.2008.00761.x.
  28. ^ Lü Junchang; Ji Qiang; Wei Xuefang; Liu Yongqing (2011). "A new ctenochasmatoid pterosaur from the Early Cretaceous Yixian Formation of western Liaoning, China". Cretaceous Research. 34: 26–30. doi:10.1016/j.cretres.2011.09.010.
  29. ^ Lü J. (2009). "A baby pterodactyloid pterosaur from the Yixian Formation of Ningcheng, Inner Mongolia, China". Acta Geologica Sinica. 83 (1): 1–8. doi:10.1111/j.1755-6724.2009.00001.x.
  30. ^ Lü, J. (2010). "A new boreopterid pterodactyloid pterosaur from the Early Cretaceous Yixian Formation of Liaoning Province, northeastern China". Acta Geologica Sinica. 24: 241–246. doi:10.1111/j.1755-6724.2010.00204.x.

Further reading

  • Barrett, P.M.; Butler, R.J.; Edwards, N.P.; Milner, A.R. (2008). Pterosaur distribution in time and space: an atlas (PDF). Flugsaurier: Pterosaur papers in honour of Peter Wellnhofer, Hone, D.W.E., and Buffetaut, E. (eds). Zitteliana B, 28. 264pp. pp. 61–107.

External links

  • The Pterosaur Database
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