Casuariiformes

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Casuariiformes
Temporal range: Miocene–present
Miocene to present Possible Paleocene appearance.
Darica Cossowary 00975.jpg
Southern cassowary
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Infraclass: Palaeognathae
Clade: Notopalaeognathae
Clade: Novaeratitae
Order: Casuariiformes
(Sclater, 1880) Forbes, 1884[1]
Families

Casuariidae
Dromaiidae

Diversity
2 families, 4 genera (including 2 extinct),
11 species (including 6 extinct)
Synonyms[2]
  • Casuarii Sclater, 1880

The Casuariiformes is an order of large flightless bird that has four surviving members: the three species of cassowary, and the only remaining species of emu. They are divided into either a single family, Casuariidae, or more typically two, with the emu splitting off into its own family, Dromaiidae.

All four living members are native to Australia-New Guinea,[3] but some possible extinct taxa occurred in other landmasses.

Systematics and evolution

Emu

The emus form a distinct family, characterized by legs adapted for running. As with all ratites, there are several contested theories concerning their evolution and relationships. As regards this order, it is especially interesting whether emus or cassowaries are the more primitive form: the latter are generally assumed to retain more plesiomorphic features, but this does not need to be true at all; the fossil record is also ambiguous, and the present state of genomics does not allow for sufficiently comprehensive analyses. A combination of all these approaches with considerations of plate tectonics at least is necessary for resolving this issue.

The total number of cassowary species described, based on minor differences in casque shape and color variations, formerly reached nine.[4] Now, however, only 3 species are recognized, and most authorities only acknowledge few subspecies or none at all.

The fossil record of casuariforms is interesting, but not very extensive. Regarding fossil species of Dromaius and Casuarius, see their genus pages.

Some Australian fossils initially believed to be from emus were recognized to represent a distinct genus, Emuarius,[5] which had a cassowary-like skull and femur and an emu-like lower leg and foot. In addition, the first fossils of mihirungs were initially believed to be from giant emus,[6] but these birds were completely unrelated.

It has been suggested that the South American genus Diogenornis was a casuraiiform bird, instead of a member of the current South American ratite lineage, the rheas. If this was the case, not only it expanded the fossil range of this lineage spatially, but temporally as well, since Diogenornis occurs in the late Paleocene and is among the earliest known ratites.[7] The Pliocene indian Hypselornis has also been linked to emus and cassowaries,[8] but no recent (post-30s) studies have been performed in this extremely obscure taxon.

Taxonomy

Casuariiformes (Sclater, 1880) Forbes 1884[9][10][11]

  • ?†Diogenornis - Alvarenga, 1983 (Paleocene of Brazil)
  • Casuariidae Kaup, 1847 [Casuariinae Reichenbach, 1849] (emus and cassowaries)
    • ?†Hypselornis Lydekker, 1929
    • Emuarius Boles, 1992 (emuwaries) (Late Oligocene – Late Miocene)
      • E. gidju (Patterson & Rich 1987) Boles, 1992 [Dromaius gidju Patterson & Rich 1987]
      • E. guljaruba Boles, 2001
    • Casuarius (Linnaeus 1758) Brisson, 1760 [Cela Oken, 1816; Cela Moehr, 1752 nomen rejectum; Rhea Lacépède, 1800 non Latham, 1790; Chelarga Billberg, 1828; Oxyporus Brookes, 1828 non (Bourdot & Galzin, 1925) Donk, 1933; Thrasys Billberg, 1828; Cassowara Perry, 1811; Hippalectryo Gloger, 1842] (cassowary)
      • C. lydekkeri Rothschild, 1911 (Pygmy cassowary)
      • C. casuarius (Linnaeus, 1758) Brisson, 1760 [Struthio casuarius Linnaeus, 1758; Casuarius casuarius altijugus (Sclater, 1878); Casuarius altijugus Sclater, 1878; ; Casuarius casuarius aruensis (Schlegel, 1866); Casuarius aruensis Schlegel, 1866; Casuarius australis D'Albertis non Wall, 1854; Casuarius casuarius australis; Casuarius casuarius beccarii (Sclater, 1875); Casuarius beccarii Sclater, 1875; Casuarius bicarunculatus Sclater, 1860; Casuarius casuarius bicarunculatus (Sclater, 1860); Casuarius bicarunculatus bicarunculatus; Casuarius bistriatus van Oort, 1907; Casuarius casuarius bistriatus van (Oort, 1907); Casuarius casuarius casuarius (Linnaeus, 1758); Casuarius casuarius chimaera Rothschild, 1904; Casuarius bicarunculatus chimaera (Rothschild, 1904); Casuarius casuarius grandis Rothschild, 1937; Casuarius galeatus Bonnaterre, 1790; Casuarius casuarius hamiltoni Mathews, 1915; Casuarius casuarius intensus Rothschild, 1898; Casuarius bicarunculatus intermedius Rothschild, 1928; Casuarius casuarius intermedius (Rothschild, 1928); Casuarius casuarius johnsonii (Müller, 1866); Casuarius johnsonii Müller, 1866; Casuarius casuarius lateralis Rothschild, 1925; Casuarius casuarius salvadorii (Oustalet, 1878); Casuarius salvadorii Oustalet, 1878; Casuarius casuarius sclaterii (Salvadori, 1878); Casuarius sclaterii Salvadori, 1878; Casuarius casuarius tricarunculatus (Beccari, 1876); Casuarius bicarunculatus tricarunculatus (Beccari, 1876); Casuarius tricarunculatus Beccari, 1876; Casuarius casuarius violicollis Rothschild, 1899; Cassowara eximia Perry, 1811; Hippalectryo indicus Gloger 1842; Hippalectryo casuarius; Casuarius hagenbecki Rothschild, 1904; Casuarius casuarius hagenbecki; Casuarius unappendiculatus hagenbecki (Rothschild, 1904); Casuarius emeu; Casuarius orientalis; Casuarius javanensis] (Southern Cassowary)
      • C. unappendiculatus Blyth 1860 [Casuarius doggetti Rothschild, 1904; ; Casuarius unappendiculatus doggetti (Rothschild, 1904); Casuarius unappendiculatus mitratus Rothschild, 1904; Casuarius unappendiculatus multicolor Le Souef, 1930; Casuarius unappendiculatus suffusus Rothschild, 1904; Casuarius unappendiculatus rothschildi (Matschie, 1901); Casuarius rothschildi Matschie, 1901; Casuarius unappendiculatus philipi (Rothschild, 1898); Casuarius philipi Rothschild, 1898; Casuarius unappendiculatus unappendiculatus; Casuarius unappendiculatus occipitalis (Salvadori, 1878); Casuarius occipitalis Salvadori, 1878; Casuarius unappendiculatus rufotinctus Rothschild, 1900; Casuarius unappendiculatus aurentiacus Rothschild, 1899; Casuarius kaupi Rosenberg, 1861; Casuarius laglaizei Oustalet, 1893] (Northern Cassowary)
      • C. bennetti Gould, 1857 [Casuarius westermanni Sclater, 1874; Casuarius papuanus Schlegel, 1871; Casuarius goodfellowi Rothschild, 1914; Casuarius foersteri Rothschild, 1913; Casuarius keysseri Rothschild, 1912; Casuarius jamrachi Rothschild, 1904; Casuarius roseigularis Rothschild, 1905; Casuarius rogersi Rothschild, 1928; Casuarius edwardsi Oustalet, 1878; Casuarius claudii Ogilvie-Grant, 1911; Casuarius picticollis Sclater, 1874; Casuarius loriae Rothschild, 1898; Casuarius maculatus Rothschild, 1900] (Dwarf Cassowary)
        • C. b. westermanni (Sclater, 1874) [Casuarius westermanni Sclater, 1874; Casuarius bennetti westermanni; Casuarius papuanus Schlegel, 1871; Casuarius bennetti papuanus (Schlegel, 1871); Casuarius goodfellowi Rothschild, 1914; Casuarius bennetti goodfellowi (Rothschild, 1914); Casuarius papuanus goodfellowi (Rothschild, 1914); Casuarius papuanus shawmayeri Rothschild, 1937; Casuarius bennetti shawmayeri (Rothschild, 1937); Casuarius foersteri Rothschild, 1913; Casuarius bennetti foersteri (Rothschild, 1913); Casuarius picticollis hecki Rothschild, 1899; Casuarius bennetti hecki (Rothschild, 1899); Casuarius keysseri Rothschild, 1912; Casuarius bennetti keysseri (Rothschild, 1912); Casuarius jamrachi Rothschild, 1904; Casuarius casuarius jamrachi (Rothschild, 1904); Casuarius unappendiculatus jamrachi (Rothschild, 1904); Casuarius roseigularis Rothschild, 1905; Casuarius bennetti roseigularis (Rothschild, 1905); Casuarius rogersi Rothschild, 1928] (Papuan dwarf cassowary)
        • C. b. bennetti Gould, 1857 [Casuarius edwardsi Oustalet, 1878; Casuarius bennetti edwardsi (Oustalet, 1878); Casuarius westermanni edwardsi (Oustalet, 1878); Casuarius claudii Ogilvie-Grant, 1911; Casuarius bennetti claudii (Ogilvie-Grant, 1911); Casuarius picticollis Sclater, 1874; Casuarius bennetti picticollis (Sclater, 1874); Casuarius loriae Rothschild, 1898; Casuarius bennetti loriae (Rothschild, 1898); Casuarius maculatus Rothschild, 1900; Casuarius bennetti maculatus (Rothschild, 1900)] (Bennett's cassowary)
  • Dromaiidae Huxley, 1868 [Dramaiinae Gray, 1870; Dramiceiidae Richmond, 1908; Dramaeidae Newton, 1896] (modern emus)
    • Dromaius Vieillot, 1816 [Tachea Fleming 1822; Emou Griffith & Pidgeon 1829; Peronista Mathews 1912; Metapteryx De Vis, 1892] (emus) (Middle Miocene – Recent)
      • D. ocypus Miller, 1963 [Casuarius ocypus (Miller, 1963)]
      • D. novaehollandiae (Latham, 1790) Vieillot 1816 [Casuarius novae-hollandiae Latham, 1790; †Dromaius gracilipes De Vis, 1892; †Dromaius patricius De Vris, 1888; Metapteryx bifrons De Vis, 1892] (emu)
        • D. n. minor Spencer, 1906 [Dromaeus minor (sic) Spencer, 1906; Dromaius novaehollandiae ater Vieillot, 1817; Dromiceius spenceri (sic) Mathews, 1912; Dromaeus bassi Legge, 1907; Dromaius ater Vieillot, 1817 nomen novum; Peronista spenceri (Mathews, 1912)] (King Island/black emu)
        • D. n. baudinianus Parker, 1984 [Dromaius baudinianus Parker 1984; Dromaius baudinianus baudinianus; Dromaius parvulus Mathews, 1901; Dromaius peroni Rothschild, 1907; Dromiceius novaehollandiae gunni Mathews, 1922; Peronista peroni (Rothschild 1907) Mathews, 1913] (Kangaroo Island/dwarf emu)
        • D. n. diemenensis (Jennings, 1827) Le Souef, 1907 [Casuarius diemenianus Jennings, 1827; Dromaeus diemenensis (Jennings, 1827) Le Souëf, 1907; Peronista diemenianus Mathews, 1927] (Tasmanian emu)
        • D. n. novaehollandiae (Latham, 1790) [Dromaius novaehollandiae rothschildi Mathews, 1912; Casuarius australis Shaw, 1792; Casuarius novaehollandiae Latham, 1790; Dromaeus irroratus Bartlett, 1859; Dromaius novaehollandiae montanus Campbell, 1939; Dromaius novaehollandiae woodwardi Mathews, 1912; Dromiceius emu Stephens, 1826; Dromiceius major Brookes, 1830; Tachea novaehollandiae; Struthio novaehollandiae] (Australian emu)

Footnotes

  1. ^ Brand, S. (2008)
  2. ^ Brodkob, Pierce (1963). "Catalogue of fossil birds 1- Archaeopterygiformes through Ardeiformes". Biological sciences, Bulletin of the Florida State Museum. 7 (4): 180–293. Retrieved 30 December 2015.
  3. ^ Clements, J (2007)
  4. ^ Lockyer, Norman (14 October 1875). Nature. 12. London, UK: Macmillan and Co. pp. 516–517.
  5. ^ From "Emu" + "Casuarius". Describer W. E. Boles commonly refers to the genus as "emuwaries" or "cassomus".
  6. ^ The vernacular name "mihirung" is derived from mihirung paringmal, which means "giant emu" in the Chaap Wuurong language
  7. ^ H. Alvarenga, Diogenornis fragilis Alvarenga, 1985, restudied: a South American ratite closely related to Casuariidae, 2010
  8. ^ Lowe, Percy Roycroft 1929. Some remarks on Hypselornis sivalensis Lydekker. Ibis. 71: (4) 571–576. (Journal Article)
  9. ^ Mikko's Phylogeny Archive [1] Haaramo, Mikko (2007). "PALEOGNATHIA- paleognathous modern birds". Retrieved 30 December 2015.
  10. ^ Paleofile.com (net, info) "Archived copy". Archived from the original on 2016-01-11. Retrieved 2015-12-30.. "Taxonomic lists- Aves". Archived from the original on 11 January 2016. Retrieved 30 December 2015.
  11. ^ [2] Perron, Richard (2010). "Taxonomy of the Genus Casuarius". Retrieved 30 December 2015.

See also

References

  • Boles, Walter E. (2001). "A new emu (Dromaiinae) from the Late Oligocene Etadunna Formation". Emu. 101: 317–321. doi:10.1071/MU00052.
  • Brands, Sheila (Apr 8, 2012). "Taxon: Order Casuariiformes". Project: The Taxonomicon. Retrieved Jun 17, 2012.
  • Clements, James (2007). The Clements Checklist of the Birds of the World (6 ed.). Ithaca, NY: Cornell University Press. ISBN 978-0-8014-4501-9.
  • Folch, A. (1992). "Family Casuariidae (Cassowaries)". In del Hoyo, Josep; Elliott, Andrew; Sargatal, Jose. Handbook of the Birds of the World, Volume 1: Ostrich to Ducks. Barcelona, Spain: Lynx Edicions. pp. 90–97. ISBN 84-87334-09-1.

External links

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