2013 in paleontology

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List of years in paleontology
In science
2010
2011
2012
2013
2014
2015
2016
Species described

Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2013.

Plants

Angiosperms

Name Novelty Status Authors Age Unit Location Notes Images

Anonaspermum warmanensis[2]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A member of Annonaceae.

Atli[3]

Gen. et sp. nov.

Valid

Smith et al.

Late Cretaceous (Campanian)

 Canada

A ranunculalean known from a stem with lianoid anatomy. The type specimen is Atli morinii.

Bambusiculmus[4]

Gen. et 2 sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo. The type specimen is Bambusiculmus angustus; genus also includes Bambusiculmus latus.

Bambusium angustifolia[4]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo, a species of Bambusium.

Bambusium latifolia[4]

Sp. nov.

Valid

Wang & Zhou in Wang et al.

Miocene

Dajie Formation

 China

A bamboo, a species of Bambusium.

Bombacacioxylon[5]

Gen. et sp. nov.

Valid

Mehrotra et al.

Late Miocene to early Pliocene

Tipam Group

 India

A possible relative of Bombax. The type specimen is Bombacacioxylon tertiarum.

Carpolithus warmanensis[2]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A fossil fruit.

Cornerocarpon[2]

Gen. et sp. nov.

Valid

Grote in Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A member of Rosales belonging or related to the family Moraceae. Genus includes new species C. copiosum.

Corylopsis grotei[2]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A species of Corylopsis.

Diospyros neyveliensis[6]

Sp. nov.

Valid

Mukherjee & Prasad

Miocene

Neyveli Formation

 India

A member of the family Ebenaceae described on the basis of fossil wood. Originally described as a species of Diospyros; Srivastava et al. (2018) transferred this species to the genus Ebenoxylon.[7]

Firmiana oligocenica[8]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Firmiana.

Grotea[2]

Gen. et sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A member of the family Bignoniaceae. Genus includes new species G. warmanensis.

Guadua morronei[9]

Sp. nov.

Valid

Brea, Zucol & Franco

Pliocene–Pleistocene

Ituzaingó Formation

 Argentina

A bamboo, a species of Guadua.

Hamamelis warmanensis[2]

Sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A witch-hazel.

Iterophyllum[10]

Gen. et sp. nov.

Valid

Barral et al.

Early Cretaceous (late Barremian)

 Spain

A basal eudicot. The type specimen is Iterophyllum lobatum.

Jaguariba wiersemana[11]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

 Brazil

Fossil Nymphaeales.

Lagerstroemia corvinusii[12]

Sp. nov.

Valid

Prasad

Mio-Pliocene

Churia Formation

   Nepal

A species of Lagerstroemia.

Laurelia otagoensis[13]

Sp. nov

Valid

Conran, Bannister & Lee

Early Miocene

 New Zealand

A species of Laurelia.

Lauroflorum[2]

Gen. et sp. nov.

Valid

Wang, Blanchard & Dilcher

Eocene

Cockfield Formation

 United States

A member of Lauraceae. Genus includes new species L. warmanensis.

Lindera acuminatissima[14]

Sp. nov

Valid

Dao & Sun in Dao et al.

Late Pliocene

Mangbang Formation

 China

A species of Lindera.

Micropetasos[15]

Gen. et sp. nov

Valid

Poinar, Chambers & Wunderlich

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

An early flowering plant of uncertain phylogenetic placement. Genus includes new species M. burmensis.

Myrtineoxylon[16]

Gen. et sp. nov.

Valid

Oskolski, Feng & Jin

Late Eocene

Youganwo Formation

 China

A member of Myrtaceae described on the basis of fossil wood. Genus includes new species M. maomingensis.

Palmoxylon vastanensis[17]

Sp. nov.

Valid

Prasad et al.

Eocene

Cambay Shale Formation

 India

A palm tree, a species of Palmoxylon.

Paranephelium makumensis[8]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Paranephelium.

Poeciloneuron preindicum[18]

Sp. nov.

Valid

Srivastava & Mehrotra

Late Oligocene

 India

A species of Poeciloneuron.

Pongamia arjunkholaensis[12]

Sp. nov.

Valid

Prasad

Mio-Pliocene

Churia Formation

   Nepal

A species of Pongamia.

Potomacapnos[19]

Gen. et sp. nov.

Valid

Jud & Hickey

Early Cretaceous (Aptian)

Potomac Group

 United States

A close relative of ranunculalean eudicots. The type specimen is Potomacapnos apeleutheron.

Pterygota palaeoalata[8]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Pterygota.

Sapindus palaeoemarginatus[8]

Sp. nov.

Valid

Srivastava & Mehrotra

Oligocene (Chattian)

Tikak Parbat Formation

 India

A species of Sapindus.

Spixiarum kipea[20]

Gen. nov. et sp. nov.

Valid

Coiffard, Mohr & Bernardes-de-Oliveira

Early Cretaceous

Crato Formation

 Brazil

Fossil Aroid.

Virola dominicana[21]

Sp. nov.

Valid

Poinar & Steeves

Cenozoic (Eocene or Miocene)

 Dominican Republic

A member of Myristicaceae found in Dominican amber, a species of Virola.

Other seed plants

Name Novelty Status Authors Age Unit Location Notes Images

Allicospermum angrenicum[22]

Sp. nov.

Valid

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

A Ginkgo-like plant described on the basis of seeds.

Bancroftiastrobus[23]

Gen. et sp. nov.

Valid

Rothwell et al.

Late Jurassic (early Kimmeridgian)

Kimmeridge Clay Formation

 United Kingdom

A member of Coniferales of uncertain phylogenetic placement. The type specimen is Bancroftiastrobus digitata.

Chengia[24]

Gen. et sp. nov.

Valid

Yang, Lin & Wang

Early Cretaceous (Aptian)

Yixian Formation

 China

A member of Ephedraceae. The type specimen is Chengia laxispicata.

Cunninghamia taylorii[25]

Sp. nov.

Valid

Serbet, Bomfleur & Rothwell

Late Cretaceous (Campanian)

Horseshoe Canyon Formation

 Canada

A species of Cunninghamia.

Donponoxylon[26]

Gen. et 2 sp. nov.

Valid

Tidwell, Britt & Wright

Middle to Late Jurassic

 Australia
 New Zealand

A seed plant of uncertain phylogenetic placement. The type specimen is Donponoxylon bennettii; genus also contains Donponoxylon jacksonii.

Dordrechtites arcanus[27]

Sp. nov.

Valid

Bergene, Taylor & Taylor

Middle Triassic (Anisian)

Fremouw Formation

 Antarctica

A gymnosperm of uncertain phylogenetic placement.

Elatocladus anorovii[28]

Sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

 Russia

A conifer of uncertain phylogenetic placement.

Friedsellowia gracilifolia[29]

Gen. nov. et sp. nov.

Valid

Löwe et al.

Early Cretaceous

Crato Formation

 Brazil

A gnetophyte. The type of a new genus.

Itajuba[30]

Gen. et sp. nov

Valid

Ricardi-Branco et al.

Early Cretaceous (Aptian or Albian)

Crato Member of the Santana Formation

 Brazil

A member of Gnetales. The type specimen is Itajuba yansanae.

Nagrenia[22]

Gen. et sp. nov.

Valid

Nosova

Middle Jurassic

Angren Formation

 Uzbekistan

A member of Ginkgoales described on the basis of peduncles furcating into several pedicels. Genus includes new species N. samylinae.

Pagiophyllum heshanense[31]

Sp. nov.

Valid

Liu & Yao

Permian

 China

A species of Pagiophyllum.

Pararaucaria delfueyoi[32]

Sp. nov.

Valid

Escapa et al.

Late Jurassic

Cañadón Calcáreo Formation

 Argentina

A member of Coniferales belonging to the family Cheirolepidiaceae, a species of Pararaucaria.

Picea yanensis[28]

Sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

 Russia

A spruce.

Prumnopitys anglica[33]

Sp. nov.

Valid

Greenwood, Hill & Conran

Eocene

Branksome Formation

 United Kingdom

A species of Prumnopitys.

Schizolepidopsis canicularis[34]

Sp. nov.

Valid

Leslie et al.

Early Cretaceous (probably Aptian-Albian)

Tevshiin Govi Formation

 Mongolia

A member of Coniferales, possibly a member of the stem group of Pinaceae; a species of Schizolepidopsis.

Stephanospermum braidwoodensis[35]

Sp. nov.

Valid

Spencer, Hilton & Sutton

Carboniferous

Mazon Creek fossil beds

 United States

A member of Medullosales, a species of Stephanospermum.

Stephanospermum shuichengensis[36]

Sp. nov.

Valid

Spencer et al.

Permian (Lopingian)

Wangjiazhai Formation

 China

A member of Medullosales, a species of Stephanospermum.

Xuanweioxylon[37]

Gen. et sp. nov.

Valid

He et al.

Permian (late Wuchiapingian-Changhsingian)

Xuanwei Formation

 China

A member of Coniferophyta. Genus includes new species X. scalariforme.

Other plants

Name Novelty Status Authors Age Unit Location Notes Images

Acrolejeunea ucrainica[38]

Sp. nov

Valid

Mamontov, Heinrichs & Schäfer-Verwimp in Mamontov et al.

Late Eocene

Rovno amber

 Ukraine

A liverwort belonging to the family Lejeuneaceae.

Cymatiosphaera scitula[39]

Sp. nov.

Valid

Wicander & Playford

Devonian (Famennian)

Saverton Shale

 United States

A member of Chlorophyta belonging to the group Prasinophyceae.

Ashicaulis beipiaoensis[40]

Sp. nov

Valid

Tian et al.

Middle Jurassic

Tiaojishan Formation

 China

A member of Osmundaceae. Originally described as a species of Ashicaulis; transferred to the genus Millerocaulis by Bomfleur, Grimm & McLoughlin (2017).[41]

Brachytheciites sachalinensis[42]

Sp. nov

Valid

Ignatov & Perkovsky

Middle Eocene

Sakhalinian amber

 Russia

A moss belonging to the family Brachytheciaceae.

Bryiidites[43]

Gen. et sp. nov.

Valid

Barclay et al.

Late Cretaceous (late Cenomanian)

Middle Dakota Formation

 United States

A member of Bryidae of uncertain phylogenetic placement. The type specimen is Bryiidites utahensis.

Bryokhutuliinia crassimarginata[44]

Sp. nov

Valid

Ignatov et al.

Late Jurassic

 Russia

A moss.

Calymperites[45]

Gen. et sp. nov

Valid

Ignatov & Perkovsky

Late Eocene

Rovno amber

 Ukraine

A moss belonging to the group Dicranales (sensu lato). Genus includes new species C. ucrainicus.

Franhueberia[46]

Gen. et sp. nov.

Valid

Hoffman & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada

A basal member of Euphyllophytina. The type specimen is Franhueberia gerriennei.

Mygdykitia[28]

Gen. et sp. nov

Valid

Golovneva & Shczepetov

Late Cretaceous (Campanian)

Ola Formation

 Russia

A fern belonging to the family Dipteridaceae. Genus includes new species M. ochotica.

Palaeosorum[47]

Gen. et sp. nov

Valid

Jacques & Zhou in Jacques, Su & Zhou

Miocene

Dajie Formation

 China

A member of Polypodiaceae. The type specimen is Palaeosorum ellipticum.

Pseudoleskeellites[42]

Gen. et sp. nov

Valid

Ignatov & Perkovsky

Middle Eocene

Sakhalinian amber

 Russia

A moss belonging to the order Hypnales. Genus includes new species P. obscurus.

Todea amissa[48]

Sp. nov.

Valid

Carvalho in Carvalho et al.

Early Eocene

 Argentina

A species of Todea.

Yavanna[49]

Gen. et sp. nov

Valid

Vera

Early Cretaceous (Aptian)

Cerro Negro Formation

 Antarctica
(Livingston Island)

A member of Cyatheales of uncertain phylogenetic placement. The type specimen is Yavanna chimaerica.

Cnidarians

Name Novelty Status Authors Age Unit Location Notes Images

Alveopora jessicae[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Alveopora.

Alveopora oliveri[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Alveopora.

Arctophyllum spitsbergensis[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian)

Treskelodden Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Cyathopsidae.

Astreopora wallaceae[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Astreopora.

Aulastraea crassicalix[52]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Aulastraea.

Barentsburgia[51]

Gen. et sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Wordiekammen Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Aulophyllidae. The type species is B. crinisphyllia.

Catenipora nishioi[53]

Sp. nov

Valid

Niko & Adachi

Silurian

Gionyama Formation

 Japan

A tabulate coral belonging to the family Halysitidae.

Caulastraea cummingae[50]

Sp. nov

Valid

Bromfield

Miocene

 Fiji
 Indonesia

A stony coral, a species of Caulastraea.

Caulastraea geoffreyi[50]

Sp. nov

Valid

Bromfield

Miocene

 Indonesia

A stony coral, a species of Caulastraea.

Cyphastrea aliceae[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral belonging to the family Faviidae.

Cyphastrea buddae[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral belonging to the family Faviidae.

Cyphastrea spinea[50]

Sp. nov

Valid

Bromfield

Miocene

 Fiji

A stony coral belonging to the family Faviidae.

Euphyllia romani[50]

Sp. nov

Valid

Bromfield

Miocene

 Indonesia

A stony coral, a species of Euphyllia.

Fedorowskites[51]

Gen. et sp. nov

Valid

Chwieduk

Permian (Wordian to Lopingian)

Kapp Starostin Formation

 Norway
 Russia

A rugose coral belonging to the order Stauriida and the family Polycoeliidae. The type species is F. spitsbergensis; genus also includes "Sochkineophyllum" zavodovskyi Sokolov (1959).[54]

Fomichevella borealis[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian)

Treskelodden Formation

 Norway

A rugose coral belonging to the order Stauriida and the family Kleopatrinidae.

Galaxea kabairaensis[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Galaxea.

Galaxea salayarensis[50]

Sp. nov

Valid

Bromfield

Pliocene

 Indonesia

A stony coral, a species of Galaxea.

Gronfjordphyllum[51]

Gen. et 2 sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Wordiekammen Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Aulophyllidae. The type species is G. minor; genus also includes G. parvum.

Halysites delicatus[53]

Sp. nov

Valid

Niko & Adachi

Silurian

Gionyama Formation

 Japan

A tabulate coral belonging to the family Halysitidae.

Halysites intricatus[53]

Sp. nov

Valid

Niko & Adachi

Silurian

Gionyama Formation

 Japan

A tabulate coral belonging to the family Halysitidae.

Halysites kyushuensis[53]

Sp. nov

Valid

Niko & Adachi

Silurian

Gionyama Formation

 Japan

A tabulate coral belonging to the family Halysitidae.

Heintzella breviseptata[51]

Sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Wordiekammen Formation

 Norway

A rugose coral belonging to the order Stauriida and the family Kleopatrinidae.

Heintzella poljarica[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian)

Treskelodden Formation

 Norway

A rugose coral belonging to the order Stauriida and the family Kleopatrinidae.

Heteropistophyllum[52]

Gen. et comb. nov

Valid

Löser, Castro & Nieto

Early Cretaceous

 Slovenia
 Spain

A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. A new genus for "Pseudopistophyllum" quinqueseptatum Turnšek & Buser (1976).

Krusenella[51]

Gen. et sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Treskelodden Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Antiphyllidae. The type species is K. pachyseptata.

Leptoria hayae[50]

Sp. nov

Valid

Bromfield

Miocene

 Fiji

A stony coral, a species of Leptoria.

Leptoria traba[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Leptoria.

Leptoseris lauensis[50]

Sp. nov

Valid

Bromfield

Miocene

Lau Group

 Fiji

A stony coral belonging to the family Agariciidae.

Linnephyllum[51]

Gen. et 2 sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Wordiekammen Formation

 Norway

A rugose coral belonging to the order Stauriida and the suborder Aulophyllina. The type species is L. spitsbergensis; genus also includes L. longiseptatum.

Madracis crassisepta[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral belonging to the family Pocilloporidae.

Madracis labrum[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral belonging to the family Pocilloporidae.

Metaulastraea elae[52]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Metaulastraea.

Mitrodendron ultimus[52]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Mitrodendron.

Montipora pandolfii[50]

Sp. nov

Valid

Bromfield

Miocene

 Indonesia

A stony coral, a species of Montipora.

Montipora teresea[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Montipora.

Montipora wilsonae[50]

Sp. nov

Valid

Bromfield

Early Pleistocene

 Indonesia

A stony coral, a species of Montipora.

Multithecopora fukugakuchiensis[55]

Sp. nov

Valid

Niko, Ibaraki & Tazawa

Early Carboniferous

Omi Limestone

 Japan

A tabulate coral belonging to the order Auloporida and the family Multithecoporidae.

Paraacanthogyra hispanica[52]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Paraacanthogyra.

Parnassomeandra steuberi[56]

Sp. nov

Valid

Löser

Cretaceous (Albian and Cenomanian[57])

 France
 Greece[57]

A stony coral belonging to the superfamily Eugyroidea and the family Eugyridae.

Platygyra renemai[50]

Sp. nov

Valid

Bromfield

Miocene

 Papua New Guinea

A stony coral, a species of Platygyra.

Pleurodendron[52]

Gen. et sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral belonging to the group Heterocoeniina and the family Elasmocoeniidae. The type species is Pleurodendron prebeticum.

Pseudotimania arctica[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian) to Permian (Sakmarian)

Treskelodden Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Bothrophyllidae.

Pseudotimania borealis[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian) to Permian (Sakmarian)

Treskelodden Formation
Wordiekammen Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Bothrophyllidae.

Pseudotimania longiseptata[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian) to Permian (Sakmarian)

Treskelodden Formation
Wordiekammen Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Bothrophyllidae.

Silingastraea valentina[52]

Sp. nov

Valid

Löser, Castro & Nieto

Early Cretaceous (Albian)

 Spain

A stony coral, a species of Silingastraea.

Symphyllia densium[50]

Sp. nov

Valid

Bromfield

Early Pleistocene

 Indonesia

A stony coral belonging to the family Mussidae.

Tschussovskenia borealis[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian)

Treskelodden Formation

 Norway

A rugose coral belonging to the order Stauriida and the family Lithostrotionidae.

?Tschussovskenia columellata[51]

Sp. nov

Valid

Chwieduk

Permian (Sakmarian)

Treskelodden Formation

 Norway

A rugose coral belonging to the order Stauriida and the family Lithostrotionidae.

Turbinaria jacksoni[50]

Sp. nov

Valid

Bromfield

Miocene

 Fiji

A stony coral, a species of Turbinaria.

Yakovleviella spitsbergensis[51]

Sp. nov

Valid

Chwieduk

Carboniferous (Gzhelian) and Permian (Sakmarian)

Wordiekammen Formation

 Norway

A rugose coral belonging to the group Stauriida and the family Aulophyllidae.

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Adlatipora[58]

Gen. et sp. nov

Valid

Gautier, Jackson & McKinney

Permian

 United States

An acanthocladiid bryozoan. The type species is Adlatipora fossulata.

Basyaylella[59]

Gen. et sp. nov

Valid

Zágoršek & Gordon

Late Tortonian

Mut Basin

 Turkey

An ascophoran bryozoan. Its type species is Basyaylella elsae.

Charixa goshouraensis[60]

Sp. nov

Valid[61]

Dick et al.

Cretaceous (late Albian or early Cenomanian)

 Japan

A cheilostome bryozoan. Originally described as a species of Charixa; Dick, Sakamoto & Komatsu (2018) transferred this species to the genus Kenocharixa.[62]

Charixa sagnieri[63]

Sp. nov

Valid

Taylor & Calzada

Early Cretaceous (Aptian)

 Spain

A member of Malacostegina belonging to the family Electridae.

Conopeum wilsoni[64]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Conopeum.

Haplostoechios[60]

Gen. et 2 sp. nov

Valid[61]

Dick et al.

Cretaceous (late Albian or early Cenomanian)

 Japan

A malacostegan. Genus contains two species: H. hayamiae and H. clusum.

Heloclema uralicum[65]

Sp. nov

Valid

Tolokonnikova

Early Carboniferous

 Russia

A species of Heloclema.

Monoceratopora whybrowi[64]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Monoceratopora.

Ostrovskia[59]

Gen. et sp. nov

Valid

Zágoršek & Gordon

Late Tortonian

Mut Basin

 Turkey

An ascophoran bryozoan. Its type species is Ostrovskia triforamina.

Poricellaria sakurkari[66]

Sp. nov

Valid

Sonar & Gaikwad

Eocene (Lutetian) to Miocene (Burdigalian)

 India

A member of Flustrina belonging to the family Poricellariidae.

Poricellaria waioriensis[66]

Sp. nov

Valid

Sonar & Gaikwad

Eocene (Lutetian) to Miocene (Burdigalian)

 India

A member of Flustrina belonging to the family Poricellariidae.

Primorella kodinkensis[65]

Sp. nov

Valid

Tolokonnikova

Early Carboniferous

 Russia

A species of Primorella.

Steginoporella chiplonkari[67]

Sp. nov

Valid

Sonar & Gaikwad

Oligocene (Chattian)

Maniyara Fort Formation

 India

A member of Flustrina belonging to the family Steginoporellidae.

Steginoporella mathuri[67]

Sp. nov

Valid

Sonar & Gaikwad

Miocene (Aquitanian)

 India

A member of Flustrina belonging to the family Steginoporellidae.

Steginoporella murachbanensis[67]

Sp. nov

Valid

Sonar & Gaikwad

Miocene (Burdigalian)

Chhasara Formation

 India

A member of Flustrina belonging to the family Steginoporellidae.

Tyloporella smithi[64]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Tyloporella.

Vincularia feddeni[66]

Sp. nov

Valid

Sonar & Gaikwad

Eocene (Lutetian) to Miocene (Burdigalian)

 India

A member of Flustrina belonging to the family Vinculariidae.

Wilbertopora cheethami[64]

Sp. nov

Valid

Di Martino & Taylor

Late Cretaceous (Late Campanian or Maastrichtian)

Simsima Formation

United Arab EmiratesOman border region

A species of Wilbertopora.

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Aegiromena (Aegiromenella)[68]

Subgen. et comb. nov

Valid

Rong et al.

Ordovician (Hirnantian and Katian) to Silurian (Aeronian)

Kosov Formation
Králův Dvůr Formation
Panghsa-pye Formation
Whittaker Formation

 Canada
 China
 Czech Republic
 Kazakhstan
 Myanmar

A member of Plectambonitoidea belonging to the family Xenambonitidae. The type species is "Schuchertella" planissima Reed (1915); the subgenus also includes Aegiromena durbenensis Nikitin et al. (1980), "Chonetoidea" obliqua Jin & Chatterton (1997), Aegiromena ultima Havlíček (1967) and Aegiromena urbana Havlíček (1967).

Anathyris helmersenioides[69]

Sp. nov

Valid

Rzhonsnitskaya in Modzalevskaya, Alvarez & Rzhonsnitskaya

Devonian (Frasnian)

Kuznetsk Basin

 Russia

A member of Athyridida belonging to the family Athyrididae.

Anathyris strelnaensis[69]

Sp. nov

Valid

Rzhonsnitskaya in Modzalevskaya, Alvarez & Rzhonsnitskaya

Devonian (Frasnian)

Kuznetsk Basin

 Russia

A member of Athyridida belonging to the family Athyrididae.

Anisopleurella asiatica[68]

Sp. nov

Valid

Rong et al.

Ordovician (Hirnantian)

 China

A member of Strophomenoidea belonging to the family Sowerbyellidae.

Arenorthis paranaensis[70]

Sp. nov

Valid

Benedetto, Halpern & Inchausti

Late Ordovician (Hirnantian)

Eusebio Ayala Formation

 Paraguay

A species of Arenorthis.

Askepasma saproconcha[71]

sp nov

Valid

Topper et al.

Lower Cambrian

Wirrapowie Limestone

 Australia

A member of Paterinata, a species of Askepasma.

Baoshanella[72]

Gen. et sp. nov

Valid

Xu

Ordovician (late Darriwilian)

Shihtien Formation

 China

A member of Dalmanelloidea belonging to the family Portranellidae. The type species is B. baoshanensis.

Biseptata[73]

Gen. et sp. nov

Valid

Madison

Ordovician

 Russia

A member of Strophomenida. The type species is Biseptata briani.

Chunanella[68]

Gen. et sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

Anji Formation

 China

A member of Enteletoidea belonging to the family Schizophoriidae. The type species is C. chunanensis.

Deliella delicatula[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of the family Craniidae.

Dolerorthis multicostellata[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Orthoidea belonging to the family Hesperorthidae.

Eopholidostrophia (Megapholidostrophia)[68]

Subgen. et sp. nov

Valid

Rong et al.

Silurian (Rhuddanian), possibly also Ordovician (Katian)

Anji Formation

 China
 United Kingdom?

A member of Strophomenoidea belonging to the family Eopholidostrophiidae. The type species is Eopholidostrophia (Megapholidostrophia) magnifica; the subgenus might also include Eopholidostrophia mediocostalis (Reed, 1905).

Eoplectodonta (Eoplectodonta) boucoti[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Strophomenoidea belonging to the family Sowerbyellidae.

Eospirifer (Eospirifer) eosinensis[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Spiriferida belonging to the family Eospiriferidae.

Eospirifer ghobadiae[74]

Sp. nov

Valid

Popov & Cocks

Silurian (Aeronian)

 Iran

A member of Spiriferida belonging to the family Eospiriferidae.

Eospirifer (Protospirifer)[68]

Subgen. nov

Valid

Rong et al.

Ordovician (Katian and Hirnantian)

Changwu Formation
Arndell Sandstone?

 China
 Australia?

A member of Spiriferida belonging to the family Eospiriferidae. The type species is Eospirifer praecursor Rong, Zhan & Han (1994).

Epitomyonia subquadrata[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Dalmanelloidea belonging to the family Dicoelosiidae.

Fardenia (Fardenia) flexa[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Orthotetida belonging to the family Chilidiopsidae.

Gemmarcula buerai[75]

Sp. nov

Valid

Calzada, Maza & Moreno

Early Cretaceous (Aptian)

 Spain

A member of the family Dallinidae.

Hesperorthis orientalis[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Orthoidea belonging to the family Hesperorthidae.

Iranospirifer[74]

Gen. et sp. nov

Valid

Popov & Cocks

Silurian (Aeronian)

Qarabil Limestone

 Iran

A member of Spiriferida belonging to the family Hedeinopsidae. The type species is I. qarabilensis.

Katastrophomena zheganensis[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Strophomenoidea belonging to the family Strophomenidae.

Leptagonia? lakhalensis[76]

Sp. nov

Valid

Popov in Ghobadi Pour et al.

Devonian (Frasnian)

 Iran

A member of Strophomenida belonging to the group Strophomenoidea and the family Rafinesquinidae.

Menathyris[77]

Gen. et sp. nov

Valid

Feldman

Middle Triassic (early Ladinian)

Saharonim Formation

 Israel

A zeilleriid terebratulid. The type species is Menathyris wilsoni.

Mendacella mutabilis[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Dalmanelloidea belonging to the family Rhipidomellidae.

Neochonetes (Huangichonetes) archboldi[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes semicircularis[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes (Sommeriella) rectangularis[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Neochonetes (Sommeriella) waterhousei[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Neochonetes.

Notoconchidium argentium[79]

Sp. nov

Valid

Wright & Garratt

Late Silurian

Florence Quartzite
Mount Ida Formation

 Australia

A member of Rhynchonellida belonging to the family Hebetoechiidae.

Notoconchidium talenti[79]

Sp. nov

Valid

Wright & Garratt

Late Silurian

Mount Ida Formation

 Australia

A member of Rhynchonellida belonging to the family Hebetoechiidae.

Ochyromena[80]

Gen. et sp. nov

Valid

Zhan et al.

Ordovician (Dapingian)

Daguanshan Formation

 China

A member of Strophomenoidea belonging to the family Strophomenidae. The type species is O. plana.

Paralenorthis costata[81]

Sp. nov

Valid

Zhan et al.

Ordovician (Darriwilian)

Chiatsun Group

 China

A member of the family Orthidae.

Plectothyrella? itacurubiensis[70]

Sp. nov

Valid

Benedetto, Halpern & Inchausti

Late Ordovician (Hirnantian)

Eusebio Ayala Formation

 Paraguay

Possibly a species of Plectothyrella.

Primotimena[80]

Gen. et sp. nov

Valid

Zhan et al.

Ordovician (Dapingian)

Daguanshan Formation

 China

A member of Strophomenoidea belonging to the family Glyptomenidae. The type species is P. globula.

Proclinorthis[82]

Gen. et sp. nov

Valid

Colmenar, Villas & Vizcaïno

Late Ordovician

Glauzy Formation

 France

A platyorthid. The type species is Proclinorthis vailhanensis.

Pseudocrania insperata[83]

Sp. nov

Valid

Bassett et al.

Middle Ordovician (Darriwilian)

Lashkarak Formation

 Iran

A member of Craniida, a species of Pseudocrania.

Pupiaoia[72]

Gen. et sp. nov

Valid

Xu

Ordovician (late Darriwilian)

Shihtien Formation

 China

A member of Orthoidea belonging to the new family Pupiaoiidae. The type species is P. pupiaoensis.

Sapelnikoviella[84]

Gen. et sp. nov

Valid

Blodgett et al.

Silurian (Ludlow)

 United States

A gypidulinid brachiopod. Genus includes new species S. santuccii.

Shuangheella[80]

Gen. et sp. nov

Valid

Zhan et al.

Ordovician (Dapingian)

Daguanshan Formation

 China

A member of Strophomenoidea belonging to the family Rafinesquinidae. The type species is S. elongata.

Sphenope[85]

Gen. et comb. et sp. nov

Valid

Vörös

Late Jurassic to Early Cretaceous (Tithonian to Berriasian)

 Hungary
 Italy

A pygopid terebratulid. A new genus for "Terebratula" misilmerensis Gemmellaro (1871); genus also contains the new species Sphenope bifida.

Sulcatospira simplex[68]

Sp. nov

Valid

Rong et al.

Silurian (Rhuddanian)

 China

A member of Atrypida belonging to the family Atrypidae.

Tethyochonetes cheni[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Tethyochonetes.

Tethyochonetes sheni[78]

Sp. nov

Valid

Zhang et al.

Late Permian (Changhsingian)

 China

A member of Rugosochonetidae, a species of Tethyochonetes.

Yuhangella[68]

Gen. et sp. nov

Valid

Rong et al.

Ordovician (Hirnantian) and Silurian (Rhuddanian)

Anji Formation
Yankou Formation

 China

A member of Dalmanelloidea belonging to the family Dalmanellidae. The type species is Y. yui.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Abertella miskellyi[86]

Sp. nov

Valid

Kroh et al.

Miocene

Camarones Formation

 Argentina

An abertellid sand dollar, a species of Abertella.

Alternacantha[87]

Gen. et 4 sp. nov

Valid

Thuy & Meyer

Jurassic (Toarcian to Kimmeridgian)[88])

Chari Formation[88]

 France[88]
 India[88]
  Switzerland
 Morocco?[88]

An ophiacanthid brittle star. The type species is Alternacantha occulta; genus also includes A. schwermannorum, A. arges and A. dilionessa.[88]

Aragocystites[89]

Gen. et sp. nov

Valid

Zamora

Middle Cambrian

Murero Formation

 Spain

An edrioasteroid. The type species is Aragocystites belli.

Bathyovulaster[90]

Gen. et sp. nov

Valid

Smith

Santonian

 Italy

A spatangoid. The type species is Bathyovulaster disjunctus.

Bellastrella[91]

Gen. et sp. nov

Valid

Müller, Hahn and Bohatý

Middle Devonian

 Germany

An agelacrinitid edrioasteroid. The type species is Bellastrella eifeliana.

Ctenocrinus branisai[92]

Sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Belèn Formation

 Bolivia

A crinoid, a species of Ctenocrinus.

Cyclolampas altus[93]

Sp. nov

Valid

Saucede et al.

Middle Jurassic (late Callovian)

 France

An echinoid, a species of Cyclolampas.

Dermacantha[88]

Gen. et 3 sp. et comb. nov

Valid

Thuy

Jurassic (Hettangian to Kimmeridgian)

 Belgium
 France
 Germany
 Luxembourg
 United Kingdom

A brittle star belonging to the family Ophiacanthidae. The type species is D. leonorae; genus also includes new species D. pattyana and D. carli, as well as "Ophiocten" seeweni Kutscher & Hary (1991).

Dermocoma faberi[88]

Sp. nov

Valid

Thuy

Early Jurassic (Hettangian)

 Belgium
 Luxembourg

A brittle star belonging to the family Ophiacanthidae.

Dermocoma longwyensis[88]

Sp. nov

Valid

Thuy

Middle Jurassic (Bajocian)

 France

A brittle star belonging to the family Ophiacanthidae.

Dermocoma numbergerorum[88]

Sp. nov

Valid

Thuy

Late Jurassic (Oxfordian)

 France

A brittle star belonging to the family Ophiacanthidae.

Dermocoma potti[88]

Sp. nov

Valid

Thuy

Early Jurassic (Pliensbachian)

 France

A brittle star belonging to the family Ophiacanthidae.

Dermocoma simonschneideri[88]

Sp. nov

Valid

Thuy

Late Jurassic (Kimmeridgian)

Amaral Formation
La Pointe du Chay

 France
 Portugal

A brittle star belonging to the family Ophiacanthidae.

Eolaxoporus[88]

Gen. et 2 sp. nov

Valid

Thuy

Triassic (Anisian and Carnian) and Early Jurassic (late Sinemurian to early Pliensbachian)

Cassian Formation
Hanwang Formation
Trochitenkalk Formation

 Austria
 China
 Germany
 Italy

A brittle star belonging to the family Ophiacanthidae. The type species is E. hagdorni; genus also includes E. imminens.

Eozonella[94]

Gen. et sp. nov

Valid

Thuy, Marty & Comment

Late Jurassic (late Oxfordian)

  Switzerland

An ophiolepidid brittle star. The type species is Eozonella bergeri.

Europacantha[88]

Gen. et sp. nov

Valid

Thuy

Middle Triassic (Anisian)

Felsöörs Formation

 Hungary

A brittle star belonging to the family Ophiacanthidae. The type species is E. paciphila.

Geromura[88]

Gen. et 2 sp. nov

Valid

Thuy

Middle Jurassic (Bajocian-Bathonian boundary) to Early Cretaceous (Valanginian)

 Austria
 France

A brittle star belonging to the family Ophiacanthidae. The type species is G. teckliformis; genus also includes G. touertensis.

Graciacystis[95]

Gen. et sp. nov

Valid

Zamora, Rahman & Smith

Middle Cambrian

 Spain

A basal cinctan echinoderm. The type species is Graciacystis ambigua.

Griphocrinus pirovanoi[92]

Sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Icla Formation

 Bolivia

A crinoid, a species of Griphocrinus.

Hanshessia[87]

Gen. et sp. nov

Valid

Thuy & Meyer

Early[88] and Middle Jurassic

Chari Formation[88]

 Austria[88]
 France[88]
 Germany[88]
 India[88]
  Switzerland

An ophiacanthid brittle star. The type species is Hanshessia trochitophila.

Inexpectacantha ritae[88]

Sp. nov

Valid

Thuy

Early Jurassic (Hettangian)

 Belgium
 Luxembourg

A brittle star belonging to the family Ophiacanthidae.

Inexpectacantha weisi[88]

Sp. nov

Valid

Thuy

Early Jurassic (Hettangian to Sinemurian)

 Belgium
 Luxembourg

A brittle star belonging to the family Ophiacanthidae.

Ishidacantha[88]

Gen. et comb. et 2 sp. nov

Valid

Thuy

Jurassic (Toarcian to Oxfordian)

Chari Formation

 France
 Germany
 India

A brittle star belonging to the family Ophiacanthidae. The type species is "Ophiopholis" trispinosa Hess (1965); genus also includes new species I. hirokoae and I. fuersichi.

Juracantha[94]

Gen. et sp. nov

Valid

Thuy, Marty & Comment

Late Jurassic (late Oxfordian)

  Switzerland

An ophiacanthid brittle star. The type species is Juracantha hottingeri.

Kosachenkoastrus[96]

Gen. et sp. nov

Valid

Rozhnov

Middle Ordovician

 Russia

A member of Parablastoidea. The type species is Kosachenkoastrus volkhovensis.

Krohcoma[88]

Gen. et 2 sp. nov

Valid

Thuy

Jurassic (late Sinemurian to Kimmeridgian)

Amaral Formation
La Pointe du Chay
Patcham Formation

 Austria
 France
 India
 Portugal

A brittle star belonging to the family Ophiacanthidae. The type species is K. mira; genus also includes K. ampla.

Lapidaster[88]

Gen. et 9 sp. et comb. nov

Valid

Thuy

Jurassic and Early Cretaceous (late Sinemurian to Albian)

Bärschwil Formation
Chari Formation

 Austria
 France
 Germany
 India
  Switzerland
 United Kingdom

A brittle star belonging to the family Ophiacanthidae. The type species is L. hystricarboris; genus also includes L. caeloscopus, L. coreytaylori, L. etteri, L. lukenederi, L. mastodon, L. mathcore, L. wolfii and L. varuna, as well as "Sinosura" fasciata Kutscher & Villier (2003).

Lutocrinus[92]

Gen. et sp. nov

Valid

Thompson, Ausich & Smith

Early Devonian (Emsian)

Icla Formation

 Bolivia

A crinoid. The type species is Lutocrinus boliviaensis.

Magnuscrinus kammeri[97]

Sp. nov

Valid

Krivicich, Ausich & Keyes

Carboniferous (Viséan)

Fort Payne Formation

 United States

A crinoid.

Manfredura[88]

Gen. et comb. nov

Valid

Thuy

Late Cretaceous (Maastrichtian)

 Germany

A brittle star belonging to the family Ophiacanthidae. The type species is "Ophiomyxa" curvata Kutscher & Jagt (2000).

Megaradialocrinus dorotheakochae[98]

Sp. nov

Valid

Bohatý & Hein

Devonian

 Germany

A camerate crinoid belonging to the group Monobathrida and the family Hexacrinitidae.

Nanocarpus guoleensis[99]

Sp. nov

Valid

Chen & Han

Cambrian (Furongian)

Guole Formation

 China

A member of Stylophora.

Ophiocamax dorotheae[88]

Sp. nov

Valid

Thuy

Late Jurassic (Oxfordian)

 France
  Switzerland

A brittle star belonging to the family Ophiacanthidae.

Ophiacantha jaegeri[88]

Sp. nov

Valid

Thuy

Early Cretaceous (Hauterivian)

 Germany

A brittle star belonging to the family Ophiacanthidae.

Ophiacantha reginae[88]

Sp. nov

Valid

Thuy

Late Cretaceous (late Campanian)

 Germany

A brittle star belonging to the family Ophiacanthidae.

Ophiacantha steffenschneideri[88]

Sp. nov

Valid

Thuy

Oligocene (Rupelian)

 Germany

A brittle star belonging to the family Ophiacanthidae.

Ophiogaleus[88]

Gen. et comb. et sp. nov

Valid

Thuy

Jurassic and Cretaceous (late Sinemurian to Maastrichtian)

Agardhfjellet Formation[100] Chari Formation

 Austria
 Belgium
 Denmark
 France
 Germany
 India
 Norway[100]
  Switzerland

A brittle star belonging to the family Ophiacanthidae. The type species is "Ophiacantha" constricta Hess (1966); genus also includes "Ophiacantha" danica Rasmussen (1952) and "Ophiacantha" dorecki Hess (1962), as well as new species O. stans.

Ophiojagtus[88]

Gen. et 2 sp. et comb. nov

Valid

Thuy

Late Jurassic and Cretaceous (Oxfordian to Maastrichtian)

Amaral Formation
La Pointe du Chay

 Belgium
 France
 Germany
 Portugal
  Switzerland
 United States

A brittle star belonging to the family Ophiacanthidae. The type species is O. acklesi; genus also includes new species O. irimurai, as well as "Hemieuryale" argoviensis Hess (1966) and "Ophiosmilax" alternatus Kutscher & Jagt (2000).

Ophioleviathan[88]

Gen. et sp. nov

Valid

Thuy

Early Jurassic (late Sinemurian to late Pliensbachian)

 Austria
  Switzerland

A brittle star belonging to the family Ophiacanthidae. The type species is O. watsoni.

Ophiolimna kucerai[88]

Sp. nov

Valid

Thuy

Early Cretaceous (latest Aptian to earliest Albian)

Atlantic Ocean
(Blake Nose)

A brittle star belonging to the family Ophiacanthidae.

Ophiolimna lisae[88]

Sp. nov

Valid

Thuy

Late Jurassic (Kimmeridgian)

Amaral Formation
La Pointe du Chay

 France
 Portugal

A brittle star belonging to the family Ophiacanthidae.

Ophiolimna malagasica[88]

Sp. nov

Valid

Thuy

Middle Jurassic (Bathonian to Callovian)

Patcham Formation

 India

A brittle star belonging to the family Ophiacanthidae.

Ophiolimna tiamatia[88]

Sp. nov

Valid

Thuy

Early Jurassic (late Sinemurian to early Pliensbachian)

 Austria

A brittle star belonging to the family Ophiacanthidae.

Ophiologimus aiodipius[88]

Sp. nov

Valid

Thuy

Early Cretaceous (latest Aptian to earliest Albian)

Atlantic Ocean
(Blake Nose)

A brittle star belonging to the family Ophiacanthidae.

Ophiologimus martynovi[88]

Sp. nov

Valid

Thuy

Early Cretaceous (latest Aptian to earliest Albian)

Atlantic Ocean
(Blake Nose)

A brittle star belonging to the family Ophiacanthidae.

Ophiomalleus[88]

Gen. et 2 sp. nov

Valid

Thuy

Middle Jurassic (Callovian) to Early Cretaceous (Hauterivian)

 Germany

A brittle star belonging to the family Ophiacanthidae. The type species is O. beneficarum; genus also includes O. stevenwilsoni.

Ophiosternle[101]

Gen. et comb. nov

Valid

Thuy & Schulz

Late Jurassic (Kimmeridgian)

Mergelstetten Formation

 Germany

A brittle star belonging to the family Ophiacanthidae. The type species is "Ophiurites" crinitus Quenstedt (1876).

Ophiotholia aurora[87]

Sp. nov

Valid

Thuy & Meyer

Middle Jurassic

  Switzerland

An ophiomycetid brittle star, a species of Ophiotholia.

Ophiotoma charlottae[88]

Sp. nov

Valid

Thuy

Late Jurassic (Kimmeridgian)

 France

A brittle star belonging to the family Ophiacanthidae.

Ophiotoma incredibilis[88]

Sp. nov

Valid

Thuy

Early Cretaceous (latest Aptian to earliest Albian)

Atlantic Ocean
(Blake Nose)

A brittle star belonging to the family Ophiacanthidae.

Ophiotoma vadosa[88]

Sp. nov

Valid

Thuy

Early Jurassic (Pliensbachian)

  Switzerland
 United Kingdom

A brittle star belonging to the family Ophiacanthidae.

Ophiotreta dendrophyllicola[88]

Sp. nov

Valid

Thuy

Paleocene (Danian)

 Denmark

A brittle star belonging to the family Ophiacanthidae.

Ophiotreta hedone[88]

Sp. nov

Valid

Thuy

Eocene (Lutetian)

 France

A brittle star belonging to the family Ophiacanthidae.

Ophiotreta stefaniae[88]

Sp. nov

Valid

Thuy

Late Jurassic (Oxfordian)

 Germany

A brittle star belonging to the family Ophiacanthidae.

Protoscutella palmeri[102]

Sp. nov

Valid

Osborn, Mooi & Ciampaglio

Eocene (Lutetian)

Santee Limestone

 United States

A sand dollar belonging to the group Scutellina and the family Protoscutellidae.

Regulaecystis testudineus[103]

Sp nov

Valid

Nardin & Bohatý

Devonian (Eifelian)

 Germany

A blastozoan belonging to the family Pleurocystitidae.

Reitneracantha[88]

Gen. et sp. nov

Valid

Thuy

Early Jurassic (late Sinemurian to early Pliensbachian)

 Austria

A brittle star belonging to the family Ophiacanthidae. The type species is R. dissidens.

Rhenopyrgus piojoensis[104]

Sp nov

Valid

Sumrall, Heredia, Rodríguez & Mestre

Silurian (lower Ludlow)

Los Espejos Formation

 Argentina

A rhenopyrgid edrioasteroid, a species of Rhenopyrgus.

Sabinacantha[88]

Gen. et sp. nov

Valid

Thuy

Late Jurassic (Oxfordian)

 Germany

A brittle star belonging to the family Ophiacanthidae. The type species is S. archetypa.

Simakocrinus[105]

Gen. et 2 sp. nov.

Valid

Prokop

Devonian (Pragian, Emsian and Eifelian)

 Czech Republic

A crinoid belonging to the group Disparida, possibly a member of the family Myelodactylidae. The type species is Simakocrinus facilis sp. nov (col.); genus also contains Simakocrinus diligens sp. nov (col.).

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Acanthodus humachensis[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acanthodus.

Acanthodus raqueli[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acanthodus.

Acodus primitivus[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Acodus.

Acuminatella[107]

Gen. et 2 sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Acuminatella acuminata; genus also contains Acuminatella angusta.

Bispathodus ultimus bartzschi[108]

Subsp nov

Valid

Kononova & Weyer

Devonian (late Famennian)

 Germany

A subspecies of Bispathodus ultimus.

Carnepigondolella eozoae[107]

Sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid, a species of Carnepigondolella.

Carnepigondolella medioconstricta[107]

Sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid, a species of Carnepigondolella.

Furnishina leei[109]

Sp. nov

Valid

Lee

Cambrian

Machari Formation

 South Korea

Histiodella wuhaiensis[110]

Sp. nov

Valid

Wang et al.

Ordovician

Lower Klimoli Formation

 China

Idiognathodus craticulinodosus[111]

Sp. nov

Valid

Ishida, Suzuki & Inada

Carboniferous (middle–late Bashkirian to early Moscovian)

Ko-yama Limestone Group

 Japan

Idiognathodus heckeli[112]

Sp. nov

Valid

Rosscoe & Barrick

Carboniferous

Exline Limestone
Hertha Limestone
Hushpuckney Shale
Lower Brush Creek Shale

 United States

Idiognathodus papulatus[112]

Sp. nov

Valid

Rosscoe & Barrick

Carboniferous

Hushpuckney Shale
Lower Brush Creek Shale

 United States

Idiognathodus pseudocarinatus[112]

Sp. nov

Valid

Rosscoe & Barrick

Carboniferous

Hushpuckney Shale
Lower Brush Creek Shale

 United States

Kallidontus gondwanicus[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Kallidontus.

Kraussodontus[107]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Kraussodontus peteri.

Mesogondolella daheshenensis[113]

Sp nov

Valid

Wang in Zhou et al.

Permian (probably Wordian)

Daheshen Formation

 China

A species of Mesogondolella.

Mesogondolella subgracilis[113]

Sp nov

Valid

Wang in Zhou et al.

Permian (probably Wordian)

Daheshen Formation

 China

A species of Mesogondolella.

Metapolygnathus mazzai[114]

Sp. nov

Valid

Karádi, Kozur & Görögt

Late Triassic (late Carnian and early Norian)

Csővár Limestone Formation

 Hungary
 Italy

Muellerilepis[115]

Nom. nov

Valid

Bardashev & Bardasheva

Middle Devonian (Givetian)

 Tajikistan

A replacement name for Muellerina Bardashev & Bardasheva (2012) (preoccupied).

Naimanodus[116]

Gen. et sp. nov

Valid

Tolmacheva

Middle Ordovician

 Kazakhstan
 Kyrgyzstan
 Russia

The type species is Naimanodus degtyarevi.

Neopolygnathus margaretae[108]

Sp nov

Valid

Kononova & Weyer

Devonian (late Famennian)

 Germany

A species of Neopolygnathus.

Notiodella[117]

Gen. et sp. nov

Valid

Aldridge et al.

Late Ordovician

Soom Shale Member, Cedarberg Formation

 South Africa

A member of Prioniodontida belonging to the family Balognathidae. The type species is Notiodella keblon.

Oncodella mostleri[114]

Sp. nov

Valid

Karádi, Kozur & Görögt

Late Triassic (early Norian)

Csővár Limestone Formation

 Hungary

Palmatolepis khaensis[118]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Palmatolepis marki[118]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Palmatolepis subperlobata lapoensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Parapetella[107]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. The type species is Parapetella prominens.

Polygnathus burretti[118]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis hanensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis namdipensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus communis phaphaensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus crassulus salapensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus extralobatus phoensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Polygnathus hojedki[119]

Sp. nov

Valid

Gholamalian et al.

Devonian

 Iran

A species of Polygnathus.

Polygnathus pseudocostatus[120]

Sp nov

Valid

Klapper & Vodrážková

Devonian

 United States

A species of Polygnathus.

Primatella[107]

Gen. et comb. et 2 sp. nov

Valid

Orchard

Late Triassic (late Carnian to early Norian)

Pardonet Formation

 Canada

A gondolellid ozarkodinid. A new genus for "Epigondolella" primitia Mosher (1970); genus also contains new species Primatella asymmetrica and Primatella conservativa.

Pseudopolygnathus granulosus laepensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Pseudopolygnathus granulosus salawinensis[118]

Subsp. nov

Valid

Savage

Late Devonian

 Thailand

Pseudopolygnathus maepoensis[118]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Quadralella[107]

Gen. et sp. nov

Valid

Orchard

Late Triassic (late Carnian)

Ludington Formation

 Canada

A gondolellid ozarkodinid. The type species is Quadralella lobata.

Siphonodella banraiensis[118]

Sp. nov

Valid

Savage

Late Devonian

 Thailand

Tilcarodus[106]

Gen. et comb. nov

Valid

Zeballo & Albanesi

Early Ordovician

Rupasca Formation

 Argentina

A new genus for "Utahconus" humahuacensis Albanesi and Aceñolaza (2005).

Trapezognathus? primitivus[121]

Sp. nov

Valid

Voldman, Albanesi & Zeballo in Voldman et al.

Early Ordovician

Acoite Formation

 Argentina

Originally described as a possible species of Trapezognathus; subsequently made the type species of a separate genus Zentagnathus.[122]

Utahconus purmamarcensis[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Utahconus scandodiformis[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Utahconus tortibasis[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Utahconus.

Variabiloconus crassus[106]

Sp nov

Valid

Zeballo & Albanesi

Late Cambrian (late Furongian) or early Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A species of Variabiloconus.

Fishes

Amphibians

Research

Newly named temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Broiliellus reiszi[124]

Sp. nov

Valid

Holmes, Berman & Anderson

Early Permian

El Cobre Canyon Formation

 United States

A dissorophid, a species of Broiliellus.

Parotosuchus ptaszynskii[125]

Sp nov

Valid

Sulej & Niedźwiedzki

Late Olenekian

 Poland

A species of Parotosuchus.

Reiszerpeton[126]

Gen. et sp. nov

Valid

Maddin et al.

Early Permian

Archer City Formation

 United States

A dissorophid. The type species is Reiszerpeton renascentis.

Scapanops[127]

Gen. et sp. nov

Valid

Schoch & Sues

Early Permian

Nocona Formation

 United States

A dissorophid. The type species is Scapanops neglecta.

Tersomius dolesensis[128]

Sp nov

Valid

Anderson & Bolt

Early Permian

 United States

A species of Tersomius.

Trimerorhachis greggi[129]

Sp nov

Valid

Milner & Schoch

Early Permian

 United States

A species of Trimerorhachis.

Newly named lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Batropetes niederkirchensis[130]

Sp. nov

Valid

Glienke

Early Permian (Rotliegend)

Saar–Nahe Basin

 Germany

A species of Batropetes.

Huskerpeton[131]

Gen. et sp. nov

Valid

Huttenlocker et al.

Early Permian

Eskridge Formation

 United States

A recumbirostran. The type species is Huskerpeton englehorni.

Proxilodon[131]

Gen. et comb. nov

Valid

Huttenlocker et al.

Early Permian

Speiser Formation

 United States

A recumbirostran, a new genus for "Euryodus" bonneri.

Newly named lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Aerugoamnis[132]

Gen. et sp. nov

Valid

Henrici, Báez & Grande

Early Eocene (Wasatchian)

Green River Formation

 United States

A relative of parsley frogs. The type species is Aerugoamnis paulus.

Eobarbourula[133]

Gen. et sp. nov

Valid

Folie et al.

Early Eocene

Cambay Shale Formation

 India

A bombinatorid. Its type species is Eobarbourula delfinoi.

Gracilibatrachus[134]

Gen. et sp. nov

Valid

Báez

Early Cretaceous (late Barremian)

 Spain

A relative of the family Pipidae. The type species is Gracilibatrachus avallei.

Iberobatrachus[134]

Gen. et sp. nov

Valid

Báez

Early Cretaceous (late Barremian)

 Spain

A close relative of the genus Discoglossus. The type species is Iberobatrachus angelae.

Ichthyosaura randeckensis[135]

Sp. nov

Valid

Schoch & Rasser

Miocene

 Germany

A newt related to the Alpine newt.

Indorana[133]

Gen. et sp. nov

Valid

Folie et al.

Early Eocene

Cambay Shale Formation

 India

A possible rhacophorid. Its type species is Indorana prasadi.

Leiopelma acricarina[136]

Sp. nov

Valid

Worthy et al.

Early Miocene

 New Zealand

A member of Leiopelmatidae.

Leiopelma miocaenale[136]

Sp. nov

Valid

Worthy et al.

Early Miocene

 New Zealand

A member of Leiopelmatidae.

Liaobatrachus zhaoi[137]

Sp. nov

Valid

Dong et al.

Early Cretaceous (Barremian)

Yixian Formation

 China

An anuran of uncertain phylogenetic placement, a species of Liaobatrachus.

Paranecturus[138]

Gen. et sp. nov

Valid

Demar

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States

A mudpuppy. The type species is Paranecturus garbanii.

Ukrainurus[139]

Gen. et sp. nov

Valid

Vasilyan et al.

Miocene

 Ukraine

A relative of cryptobranchids. The type species is Ukrainurus hypsognathus.

Wesserpeton[140]

Gen. et sp. nov

Valid

Sweetman & Gardner

Barremian

Wessex Formation

 England

An albanerpetontid.

Turtles

Research

  • A study on the anatomy of the brain and inner ear of the Jurassic turtle Plesiochelys etalloni is published by Paulina Carabajal et al. (2013).[141]

Newly named turtles

Name Novelty Status Authors Age Unit Location Notes Images

Adocus inexpectatus[142]

Sp. nov

Valid

Danilov et al.

Late Eocene

Youganwo Formation

 China

An adocid, a species of Adocus.

Atoposemys[143]

Gen. et sp. nov

Valid

Hutchison

Early Paleocene (Puercan)

Fort Union Formation

 United States

A trionychid. The type species is Atoposemys entopteros.

Bairdemys healeyorum[144]

Sp. nov

Valid

Weems & Knight

Late Oligocene

Chandler Bridge Formation

 United States

A podocnemidid pleurodiran, a species of Bairdemys.

Basilemys gaffneyi[145]

Sp. nov

Valid

Sullivan, Jasinski & Lucas

Late Campanian

 United States

A nanhsiungchelyid turtle, a species of Basilemys.

Brachyopsemys[146]

Gen. et sp. nov

Valid

Tong & Meylan

Early Paleocene

Ouled Abdoun Basin

 Morocco

A cryptodiran closely related to Sandownia. The type species is Brachyopsemys tingitana.

Camerochelys[147]

Gen. et sp. nov

Valid

Pérez-García & Murelaga

Early Cretaceous (Hauterivian or Barremian)

Cameros Basin

 Spain

A member of Pan-Cryptodira (the clade containing living cryptodirans and all turtles sharing a more recent common ancestor with them than with pleurodirans), possibly a xinjiangchelyid. The type species is Camerochelys vilanovai.

Cardichelyon[143]

Gen. et sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

A platysternid. The type species is Cardichelyon rogerwoodi.

?Centrochelys marocana[148]

Sp. nov

Valid

Gmira et al.

Pliocene

 Morocco

A tortoise, possibly a species of Centrochelys.

Changmachelys[149]

Gen. et sp. nov

Valid

Brinkman et al.

Early Cretaceous

Xiagou Formation

 China

A member of (likely paraphyletic) group "Macrobaenidae". The type species is Changmachelys bohlini.

Chelodina (Chelodina) murrayi[150]

Sp. nov

Valid

Yates

Late Miocene

Waite Formation

 Australia

A chelid, a species of Chelodina.

Chelonoidis lutzae[151]

Sp. nov

Valid

Zacarías et al.

Late Pleistocene (58–22 ka)

 Argentina

A tortoise, a species of Chelonoidis.

Chrysemys isoni[152]

Sp. nov

Valid

Weems & George

Miocene

Calvert Formation

 United States

A relative of the painted turtle.

Cuora chiangmuanensis[153]

Sp. nov

Valid

Naksri et al.

Late Middle or early Late Miocene

 Thailand

An Asian box turtle.

Floridemys hurdi[152]

Sp. nov

Valid

Weems & George

Miocene

Calvert Formation

 United States

A tortoise, a species of Floridemys.

Kappachelys[154]

Gen. et sp. nov

Valid

Hirayama, Isaji & Hibino

Early Cretaceous

Akaiwa Formation

 Japan

A turtle of uncertain phylogenetic placement. Originally considered to be a relative of trionychids; Skutschas et al. (2017) considered it to be either a pan-trionychian (sister taxon to Trionychia) or a pan-carettochelyid.[155] The type species is Kappachelys okurai.

Laganemys[156]

Gen. et sp. nov

Disputed

Sereno & ElShafie

Aptian or Albian

Elrhaz Formation

 Niger

An araripemydid pleurodiran. The type species is Laganemys tenerensis. Pérez-García (2018) considered this species to be a junior synonym of Taquetochelys decorata.[157]

Neochelys liriae[158]

Sp. nov

Valid

Pérez-García & de Lapparent de Broin

Eocene (Ypresian)

 France

A podocnemidid, a species of Neochelys.

Neurankylus lithographicus[159]

Sp. nov

Valid

Larson et al.

Santonian

Milk River Formation

 Canada

A neurankyline baenid, a species of Neurankylus.

Nostimochelone[160]

Gen. et sp. nov

Valid

Georgalis et al.

Early Miocene

Zeugostasion Formation

 Greece

A pleurodiran, a member of Podocnemidoidea. The type species is Nostimochelone lampra.

Ocadia tanegashimensis[161]

Sp. nov

Valid

Takahashi et al.

Early middle Miocene

Kawachi Formation

 Japan

A geoemydid, a species of Ocadia.

Ocepechelon[162]

Gen. et sp. nov

Valid

Bardet et al.

Late Cretaceous (late Maastrichtian)

Oulad Abdoun Basin

 Morocco

A relative of dermochelyids and protostegids. The type species is Ocepechelon bouyai.

Paramongolemys[163]

Gen. et sp. nov

Valid

Danilov & Sukhanov

Late Paleocene

Naranbulak Formation

 Mongolia

A basal member of Testudinoidea. The type species is Paramongolemys khosatzkyi.

Planetochelys dithyros[143]

Sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

A relative of trionychids, a species of Planetochelys.

Psilosemys[143]

Gen. et sp. nov

Valid

Hutchison

Early Eocene (Wasatchian)

 United States

The earliest North American emydid. The type species is Psilosemys wyomingensis.

Scabremys[145]

Gen. et comb. nov

Valid

Sullivan, Jasinski & Lucas

Late Campanian

 United States

A baenid cryptodiran, a new genus for "Baena" ornata Gilmore (1935).

Shandongemys[164]

Gen. et sp. nov

Valid

Li et al.

Late Cretaceous

 China

A lindholmemydid testudinoid. The type species is Shandongemys dongwuica.

Sinemys brevispinus[165]

Sp. nov

Valid

Tong & Brinkman

Early Cretaceous

Luohandong Formation

 China

A sinemydid cryptodiran, a species of Sinemys.

Spoochelys[166]

Gen. et sp. nov

Valid

Smith & Kear

Early or middle Albian

Griman Creek Formation

 Australia

A meiolaniid-like turtle. The type species is Spoochelys ormondea.

Testudo oughlamensis[148]

Sp. nov

Valid

Gmira et al.

Pliocene

 Morocco

A tortoise, a species of Testudo.

Trapalcochelys[167]

Gen. et sp. nov

Valid

Sterli, de la Fuente & Cerda

Late Cretaceous (late Campanian to early Maastrichtian)

Allen Formation

 Argentina

A relative of meiolaniids. The type species is Trapalcochelys sulcata.

Tullochelys[143]

Gen. et sp. nov

Valid

Hutchison

Maastrichtian to early Paleocene (Puercan)

Fort Union Formation
Hell Creek Formation

 United States

A chelydrid. The type species is Tullochelys montanus.

Xenochelys floridensis[168]

Sp. nov

Valid

Bourque

Late Oligocene (early Arikareean)

 United States

A kinosternid, a species of Xenochelys.

Xinjiangchelys radiplicatoides[169]

Sp. nov

Valid

Brinkman et al.

Late Middle or early Late Jurassic

Shishugou Formation

 China

A species of Xinjiangchelys.

Xinjiangchelys wusu[170]

Sp. nov

Valid

Rabi et al.

Middle Jurassic

?Qigu Formation

 China

A species of Xinjiangchelys.

Thalattosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Concavispina[171]

Gen. et sp. nov

Valid

Zhao et al.

Triassic

Xiaowa Formation

 China

A thalattosaurid thalattosaur. The type species is Concavispina biseridens

Ichthyopterygians

Name Novelty Status Authors Age Unit Location Notes Images

Chaohusaurus zhangjiawanensis[172]

Sp. nov

Valid

Chen et al.

Early Triassic

Jialingjiang Formation

 China

A species of Chaohusaurus.

Gulosaurus[173]

Gen. et sp. nov

Valid

Cuthbertson, Russell & Anderson

Early Triassic

Sulphur Mountain Formation

 Canada

A grippidian. The type species is Gulosaurus helmi.

Malawania[174]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (late Hauterivian to Barremian)

 Iraq

A basal member of Thunnosauria. The type species is Malawania anachronus.

Qianichthyosaurus xingyiensis[175]

Sp. nov

Valid

Ji, Jiang & Motani in Yang et al.

Middle Triassic (Ladinian)

Falang Formation

 China

Thalattoarchon[176]

Gen. et sp. nov

Valid

Fröbisch et al.

Middle Triassic

 United States

A merriamosaur ichthyosaur. The type species is Thalattoarchon saurophagis

Lepidosauromorphs

Newly named sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Cryonectes[177]

Gen. et sp. nov

Valid

Vincent, Bardet & Mattioli

Pliensbachian

 France

A pliosaurid. The type species is Cryonectes neustriacus

Gronausaurus[178]

Gen. et sp. nov

Disputed

Hampe

Early Cretaceous (Berriasian)

 Germany

A member of Plesiosauroidea of uncertain phylogenetic placement,[179] possibly a leptocleidid[178] or an elasmosaurid.[180] The type species is Gronausaurus wegneri. Sachs, Hornung & Kear (2016) considered it to be a junior synonym of Brancasaurus brancai.[179]

Megacephalosaurus[181]

Gen. et sp. nov

Valid

Schumacher, Carpenter & Everhart

Late Cretaceous (middle Turonian)

 United States

A pliosaurid. The type species is Megacephalosaurus eulerti.

Palatodonta[182]

Gen. et sp. nov

Valid

Neenan, Klein & Scheyer

Early Middle Triassic

 Netherlands

A relative of placodonts. The type species is Palatodonta bleekeri.

Pliosaurus carpenteri[180]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Pliosaurus kevani[180]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Pliosaurus westburyensis[180]

Sp. nov

Valid

Benson et al.

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A pliosaurid, a species of Pliosaurus

Serpianosaurus germanicus[183]

Sp. nov

Valid

Diedrich

Middle Triassic (Anisian)

 Germany

A pachypleurosaur, a species of Serpianosaurus.

Newly named rhynchocephalians

Name Novelty Status Authors Age Unit Location Notes Images

Sphenotitan[184]

Gen. et sp. nov

Valid

Martínez et al.

Late Triassic

Quebrada del Barro Formation

 Argentina

An opisthodontian rhynchocephalian. The type species is Sphenotitan leyesi.

Newly named lizards

Name Novelty Status Authors Age Unit Location Notes Images

Apsgnathus[185]

Gen. et sp. nov

Valid

Nydam, Rowe & Cifelli

Late Cretaceous (late Campanian, Judithian)

Aguja Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement. The type species is Apsgnathus triptodon.

Arcanosaurus[186]

Gen. et sp. nov

Valid

Houssaye et al.

Early Cretaceous (Barremian or Aptian)

Cameros Basin

 Spain

An anguimorph lizard, possibly a varanoid. The type species is Arcanosaurus ibericus.

Arpadosaurus sepulchralis[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Arpadosaurus.

Barbaturex[188]

Gen. et sp. nov

Valid

Head et al.

Late middle Eocene (37.2 ± 1.3 Ma)

Pondaung Formation

 Myanmar

An iguanian lizard related to the genus Uromastyx. The type species is Barbaturex morrisoni.

Bicuspidon smikros[189]

Sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Polyglyphanodontinae, a species of Bicuspidon.

Blanosaurus[190]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Early Eocene

 Belgium
 France

An amphisbaenian. The type species is Blanosaurus primeocaenus.

Camptognathosaurus[190]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Late Paleocene

 France

An amphisbaenian. The type species is Camptognathosaurus parisiensis.

Catactegenys[185]

Gen. et sp. nov

Valid

Nydam, Rowe & Cifelli

Late Cretaceous (late Campanian, Judithian)

Aguja Formation

 United States

A night lizard. The type species is Catactegenys solaster.

Cemeterius[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A platynotan lizard of uncertain phylogenetic placement. The type species is Cemeterius monstrosus.

Cnodontosaurus[189]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Anguimorpha related to xenosaurids. The type species is Cnodontosaurus suchockii.

Dakotasaurus[189]

Gen. et sp. nov

Junior homonym

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[189] The type species is Dakotasaurus gillettorum. The generic name turned out to be preoccupied by Dakotasaurus Branson & Mehl (1932); the lizard genus was subsequently renamed Dakotaseps.[193]

Desertiguana[194]

Gen. et sp. nov

Valid

Alifanov

Late Cretaceous

 Mongolia

A phrynosomatid lizard. The type species is Desertiguana gobiensis.

Distortodon[195]

Gen. et sp. nov

Valid

Makádi

Late Cretaceous (Santonian)

 Hungary

A member of Polyglyphanodontinae. The type species is Distortodon rhomboideus.

Entomophontes[187]

Gen. et 2 sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A relative of species belonging to the genus Xenosaurus. Genus contains two species: Entomophontes incrustatus and Entomophontes hutchisoni.

Eonatator coellensis[196]

Sp. nov

Valid

Páramo

Campanian

 Colombia

A mosasaur, a new species of Eonatator.

Glyptosaurus agmodon[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Glyptosaurus.

Heterodontagama[197]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Priscagamidae. The type species is Heterodontagama borsukae.

Indiagama[197]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Agamidae. The type species is Indiagama gujarata.

Lamiasaura[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A squamate of uncertain phylogenetic placement. The type species is Lamiasaura ferox.

Lonchisaurus[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A scincomorph lizard of uncertain phylogenetic placement. The type species is Lonchisaurus trichurus.

Monocnemodon[189]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Santonian)

Straight Cliffs Formation

 United States

A member of Scincomorpha of uncertain phylogenetic placement, assigned by Nydam (2013) to an informal paramacellodid-cordylid grade.[189] The type species is Monocnemodon syphakos.

Obamadon[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Hell Creek Formation
Lance Formation

 United States

A polyglyphanodontian lizard of uncertain phylogenetic placement. The type species is Obamadon gracilis.

Odaxosaurus roosevelti[189]

Sp. nov

Valid

Nydam

Late Cretaceous

Kaiparowits Formation

 United States

A member of Anguidae, a species of Odaxosaurus.

Palaeoxantusia amara[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A night lizard, a species of Palaeoxantusia.

Palepidophyma[187]

Gen. et 2 sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A night lizard. Genus contains two species: Palepidophyma paradisa and Palepidophyma lilliputiana.

Parasaniwa cynochoros[189]

Sp. nov

Valid

Nydam

Late Cretaceous

Kaiparowits Formation

 United States

A member of Platynota of uncertain phylogenetic placement, a species of Parasaniwa.

Pariguana[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

An iguanid lizard of uncertain phylogenetic placement. The type species is Pariguana lancensis.

Pelsochamops[198]

Gen. et sp. nov

Valid

Makádi

Late Cretaceous (Santonian)

Csehbánya Formation

 Hungary

A chamopsiid. The type species is Pelsochamops infrequens.

Plesiolacerta eratosthenesi[199]

Sp. nov

Valid

Čerňanský & Augé

Late Oligocene

 Germany

A lacertid lizard, a species of Plesiolacerta.

Polyodontobaena[190]

Gen. et sp. nov

Valid

Folie, Smith & Smith

Early Paleocene

Hainin Formation

 Belgium

An amphisbaenian. The type species is Polyodontobaena belgica.

Provaranosaurus fatuus[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A relative of the Chinese crocodile lizard, a species of Provaranosaurus.

Romeosaurus[200]

Gen. et 2 sp. nov

Valid

Palci, Caldwell & Papazzoni

Late Cretaceous (early Turonian to early Santonian)

 Italy

A mosasaur related to Russellosaurus and Yaguarasaurus. Genus contains two species: Romeosaurus fumanensis and Romeosaurus sorbinii.

Schillerosaurus[201]

Nom. nov

Valid

Nydam, Chure & Evans

Late Jurassic

Morrison Formation

 United States

A replacement name for Schilleria Evans & Chure (1999).

Scincoideus grassator[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Lacertiformes, a species of Scincoideus.

Socognathus brachyodon[191][192]

Sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Lance Formation

 United States

A chamopsiid polyglyphanodontian lizard, a species of Socognathus.

Suratagama[197]

Gen. et sp. nov

Valid

Rana et al.

Early Eocene (early to middle Ypresian)

Cambay Shale Formation

 India

A member of Agamidae. The type species is Suratagama neeraae.

Suzanniwana revenanta[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Pleurodonta, a species of Suzanniwana.

Tiliqua laticephala[202]

Sp. nov

Valid

Čerňanský & Hutchinson

Pliocene

 Australia

A blue-tongued skink.

Utahgenys antongai[189]

Sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A contogeniid scincomorph, a species of Utahgenys.

Webbsaurus[189]

Gen. et sp. nov

Valid

Nydam

Late Cretaceous (Cenomanian)

Dakota Formation

 United States

A scincomorph related to skinks. The type species is Webbsaurus lofgreni.

Xestops savagei[187]

Sp. nov

Valid

Smith & Gauthier

Early Eocene

Wasatch Formation

 United States

A member of Glyptosaurinae, a species of Xestops.

Newly named snakes

Name Novelty Status Authors Age Unit Location Notes Images

Cerberophis[191][192]

Gen. et sp. nov

Valid

Longrich, Bhullar & Gauthier

Late Cretaceous (late Maastrichtian)

Hell Creek Formation

 United States

An alethinophidian snake of uncertain phylogenetic placement. The type species is Cerberophis robustus.

Kataria[203]

Gen. et sp. nov

Valid

Scanferla et al.

Early Paleocene (Danian)

Santa Lucía Formation

 Bolivia

A macrostomatan snake related to caenophidians. The type species is Kataria anisodonta.

Nidophis[204]

Gen. et sp. nov

Valid

Vasile, Csiki-Sava & Venczel

Late Cretaceous (Maastrichtian)

Haţeg Basin

 Romania

A madtsoiid. The type species is Nidophis insularis.

Archosauromorphs

Newly named basal archosauromorphs

Name Novelty Status Authors Age Unit Location Notes Images

Ankylosuchus[205]

Gen. et sp. nov

Valid

Lucas, Spielmann & Hunt

Late Triassic

Colorado City Formation

 United States

A doswelliid. The type species is Ankylosuchus chinlegroupensis.

Asperoris[206]

Gen. et sp. nov

Valid

Nesbitt, Butler & Gower

Middle Triassic (probably late Anisian)

Manda beds

 Tanzania

A non-archosaurian archosauriform of uncertain phylogenetic placement. The type species is Asperoris mnyama.

Fuyuansaurus[207]

Gen. et sp. nov

Valid

Fraser, Rieppel & Chun

Middle Triassic (Ladinian)

Falang Formation

 China

A protorosaur. The type species is Fuyuansaurus acutirostris.

Jaxtasuchus[208]

Gen. et sp. nov

Valid[209]

Schoch & Sues

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A doswelliid. The type species is Jaxtasuchus salomoni.

Machaeroprosopus lottorum[210]

Sp. nov

Valid

Hungerbühler et al.

Late Triassic

Cooper Canyon Formation

 United States

A phytosaur, a species of Machaeroprosopus.

Macrocnemus obristi[211]

Sp. nov

Valid

Fraser & Furrer

Middle Triassic (Ladinian)

Prosanto Formation

  Switzerland

A protorosaur, a species of Macrocnemus.

Wannia[212]

Gen. et comb. nov

Valid

Stocker

Late Triassic

 United States

A phytosaur; a new genus for "Paleorhinus" scurriensis Langston (1949).

Archosaurs

Other reptiles

Name Novelty Status Authors Age Unit Location Notes Images

Dolerosaurus[213]

Gen. et comb. nov

Valid

Butler

Late Triassic (late Carnian)

Lunz Formation

 Austria

A diapsid of uncertain phylogenetic placement, a new genus for "Francosuchus" trauthi Huene (1939).

Ruhuhuaria[214]

Gen. et sp. nov

Valid

Tsuji, Sobral & Müller

Middle Triassic

Manda Formation

 Tanzania

A procolophonoid parareptile, possibly an owenettid. The type species is Ruhuhuaria reiszi.

Synapsids

Non-mammalian synapsids

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Diegocanis[216]

Gen. et sp. nov

Valid

Martínez, Fernandez & Alcober

Late Triassic

Ischigualasto Formation

 Argentina

A member of Eucynodontia closely related to Ecteninion and Trucidocynodon. The type species is Diegocanis elegans.

Mandagomphodon[217]

Gen. et comb. nov

Valid[218]

Hopson

Middle Triassic

Manda Formation

 Tanzania

A traversodontid cynodont, a new genus for "Scalenodon" hirschsoni Crompton (1972).

Niassodon[219]

Gen. et sp. nov

Valid

Castanhinha et al.

Late Permian

 Mozambique

A kingoriid emydopoid dicynodont. The type species is Niassodon mfumukasi.

Tambacarnifex[220]

Gen. et sp. nov

Valid[218]

Berman et al.

Early Permian

Tambach Formation

 Germany

A varanodontine varanopid. The type species is Tambacarnifex unguifalcatus.

Mammals

Other animals

Name Novelty Status Authors Age Unit Location Notes Images

Alaconcha[221]

Gen. et sp. nov

Valid

Devaere, Clausen & Steiner in Devaere et al.

Cambrian (Tommotian)

Heraultia Limestone

 France

An animal of uncertain phylogenetic placement, a low cap-shaped bilaterally symmetrical conch. The type species is A. rugosa.

Anomalocaris kunmingensis[222]

Sp. nov

Valid

Wang, Huang & Hu

Cambrian

Wulongqing Formation

 China

A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods), a species of Anomalocaris.

Cambrocornulitus[223]

Gen. et sp. nov

Valid

Yang et al.

Early Cambrian

Qiongzhusi Formation

 China

An animal of uncertain phylogenetic placement, possibly a member of Cornulitida. The type species is Cambrocornulitus rarus.

Cephalodiscus? nusplingensis[224]

Sp. nov

Valid

Schweigert & Dietl

Late Jurassic

Nusplingen Limestone

 Germany

Possibly a pterobranch belonging to the genus Cephalodiscus.

Chengjiangocaris kunmingensis[225]

Sp. nov

Valid

Yang et al.

Cambrian Stage 3

Hongjingshao Formation

 China

A stem-arthropod.

Conularia porcella[226]

Sp. nov

Valid

Robson & Young

Late Ordovician (middle Katian)

Red River Formation

 Canada

A conulariid, a species of Conularia.

Fuxianhuia xiaoshibaensis[225]

Sp. nov

Valid

Yang et al.

Cambrian Stage 3

Hongjingshao Formation

 China

A stem-arthropod.

Gracilitheca astronauta[227]

Sp. nov

Valid

Valent & Fatka

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.

Gracilitheca mirabilis[228]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Gracilitheca.

Gracilitheca triangularis[228]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Gracilitheca.

Hunanoscolex[229]

Gen. et sp. nov

Valid

Duan & Dong

Cambrian (Paibian)

Bitiao Formation

 China

A palaeoscolecid. The type species is H. campus.

Microconchus hintonensis[230]

Sp. nov

Valid

Zatoń & Peck

Carboniferous (late Mississippian)

Bluestone Formation
Hinton Formation

 United States

A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs).

Microconchus utahensis[231]

Sp. nov

Valid

Zatoń, Taylor & Vinn

Early Triassic

Virgin Formation

 United States

A member of Microconchida (a group of animals of uncertain phylogenetic placement, possibly molluscs), a species of Microconchus.

Nephrotheca betula[228]

Sp. nov

Valid

Valent, Fatka & Marek

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Nephrotheca.

Nephrotheca sophia[227]

Sp. nov

Valid

Valent & Fatka

Cambrian

Buchava Formation

 Czech Republic

A member of Hyolitha, a species of Nephrotheca.

Oktavites kemanensis[232]

Sp. nov

Valid

Zhang et al.

Silurian (Telychian)

Manbo Formation

 China

A graptolite.

Ornatoscolex[229]

Gen. et sp. nov

Valid

Duan & Dong

Cambrian (Paibian)

Bitiao Formation

 China

A palaeoscolecid. The type species is O. hunanensis.

Paraconularia bachmanni[233]

Sp. nov

Valid

Barth et al.

Late Triassic

 Germany

A conulariid, a species of Paraconularia.

Paranomalocaris[222]

Gen. et sp. nov

Valid

Wang, Huang & Hu

Cambrian

Wulongqing Formation

 China

A member of Anomalocarididae (a group of animals with uncertain phylogenetic placement, possibly stem-arthropods). The type species is Paranomalocaris multisegmentalis.

Potamophloios canadensis[234]

Sp. nov

Valid

Pisera, Siver & Wolfe

Eocene (Lutetian)

 Canada

A sponge belonging to the family Potamolepidae.

Quasiaulacera[235]

Gen. et 2 sp. nov

Valid

Copper, Stock & Jin

Late Ordovician (Hirnantian)

Ellis Bay Formation

 Canada

A stromatoporoid. Genus contains two species: Quasiaulacera stellata and Q. occidua.

Spinitheca[236]

Gen. et sp. nov

Valid

Novozhilova

Early Cambrian (Atdabanian)

 Russia

A member of Hyolitha (a group of animals of uncertain phylogenetic placement, possibly molluscs). The type species is Spinitheca sysoievi.

Wronascolex iacoborum[237]

Sp. nov

Valid

García-Bellido, Paterson & Edgecombe

Cambrian Stage 4

Emu Bay Shale

 Australia

A palaeoscolecid.

Other organisms

Name Novelty Status Authors Age Unit Location Notes Images

Gorgonisphaeridium savertonense[39]

Sp. nov.

Valid

Wicander & Playford

Devonian (Famennian)

Saverton Shale

 United States

An acritarch.

Honeggeriella[238]

Gen. et sp. nov

Valid

Matsunaga, Stockey & Tomescu

Early Cretaceous (Valanginian-Hauterivian boundary)

Longarm Formation

 Canada

A heteromerous lichen. The type species is Honeggeriella complexa.

Plumalina brevis[239]

Sp. nov

Valid

Muscente & Allmon

Devonian (Frasnian)

Ithaca Formation

 United States

An organism of uncertain phylogenetic placement, argued by different authors to be a hydrozoan[239] or a Caulerpa-like alga.[240]

Plumalina tenera[239]

Sp. nov

Valid

Muscente & Allmon

Silurian (Wenlock)

Rochester Shale

 United States

An organism of uncertain phylogenetic placement, argued by different authors to be a hydrozoan[239] or a Caulerpa-like alga.[240]

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